Part 2 (3.307Mb)

Advisory Committee on Fishery Management

REPORT OF THE

ICES CM 1997/Assess:16

MULTISPECIES ASSESSMENT WORKING GROUP

ICES Headquarters 11-19 August 1997

PART 2 OF 2

This report is not to be quoted without prior consultation with the General Secretary. The document is a report of an expert group under the auspices of the International Council for the Exploration of the Sea and does not necessarily represent the views of the Council.

International Council for the Exploration of the Sea Conseil International pour l'Exploration de la Mer

Palægade 2-4 DK-1261 Copenhagen K Denmark

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Table 3.2.2.2.3-1 Frequencies of occurrence of various magnitudes of differences in suitabilities between runs using only 1981 stomach data and only 1991 stomach data. Four replicated predators presented separate ly.

MAGNITUDE OF COD WHITING SAITHE HADDOCK

CHANGEIN SUITABILITY

>-.4 l 2 11 15

-.2 to -.4 5 15 45 34

-.l to -.2 25 32 40 28

-.05 to -.l 40 27 15 7

-.01 to -.05 85 64 36 lO

-.005 to -.01 50 19 27 2

-.001 to -.005 72 20 8 l

O to -.001 40 12 o l

O to +.001 20 4 o 7

+.001 to .005 31 13 lO 6

+.005 to .01 50 33 lO 9

+.01 to .05 108 81 29 6

+.05 to .1 57 46 19 14

+.1 to .2 27 19 16 26

+.2 to .4 9 7 3 9

>.4 o ^{2 8 2}

Table 3.2.2.2.3-2 Frequencies of occurrence of various magnitudes of differences in m2 between runs using only 1981 stomach data and on!y 1991 stomach data. Four replicated predators presented separate ly.

MAGNITUDE OF COD WHITING SAITHE HADDOCK

CHANGEINM2

>-.2 o ⁶ o o

-.1 to -.2 2 10 o o

-.05 to -.1 9 15 3 4

-.01 to -.05 64 26 14 9

-.005 to -.01 45 26 14 9

-.001 to -.005 101 42 53 16

O to -.001 201 170 97 65

O to +.001 143 42 50 52

+.001 to .005 55 25 29 6

+.005 to .01 6 10 6 o

+.01 to .05 17 4 8 l

+.05 to .l 2 lO 6 o

+.l to .2 l 4 l o

>+.2 l o o o

E:\ACFM\MA WG97\T -32ll.DOC l 0/09/97

Table 3.2.2.2.3-3 1981 run vs. 1991 run. Linear models fitting the change in suitability and M2 to the change in predator biomass. Models with both overall slope and with separate slopes for each predator are fit. Where significant fits were found, tab le en tri es are percent of variance explained, and, for separate slopes model, the species with significant parameter estimates. Changes are expressed as both absolute value (tonnes) and percent of mean biomass of the two runs.

Change in: su it M2

Predator biomass NO .015 (+)

Prey biomass NO .013 (-)

% Predator biomass NO .014 (+)

% Prey Biomass .012 (+) .023 ( -)

Separate slopes:

Predator biomass NO .072 (+ Whiting)

Prey biomass .041 (+ Sandeel) .038 (- Herring)

% Predator biomass NO .071 (+ Whiting)

% Prey biomass .043 (+ Sandeel) .058 (- Herring)

Fixed effects mod el: .250 .176

Quarter NO .025 ( + Fourth quarter)

Predator species .012 (- Whiting .016 (+ Haddock)

- Saithe)

Prey species .020 (- Whiting NO

- Norway Pout

l

+ Haddock)

Predator * Prey .137 NO

Quarter * ^{Predator .060} .0657 (-Whiting, 2nd quarter)

Quarter * ^{Prey NO NO}

Table 3.2.3.1 Frequencies of occurrence of various magnitudes of differences in suitabilities and M2 between KEYRUN and run with only five traditional predators. Frequencies for all predators combined.

MAGNITUDE OF FREQUENCY OF MAGNITUDE OF FREQUENCY OF

CHANGEIN OCCURRENCE CHANGEINM2 OCCURRENCE

SUITABILITY

> -.20 5 >-.50 l

--.10 to -.20 lO -.20 to -.5- o

-.05 to -.10 l -.10 to -.20 4

-.01 to -.05 4 -.05 to -.10 16

-.005 to -.01 11 -.01 to -.05 58

-.001 to -.005 16 -.005 to -.01 40

O to -.001 24 -.0001 to -.005 119

O. to .001 62 O to -.0001 254

+.001 to .005 87 +0 to .0001 101

+.005 to .01 171 +.0001 to .005 411

+.01 to .05 473 +.005 to .001 67

+.05 to .l 242 +.001 to .05 118

+.1 to .2 143 +.05to.1 33

+.2 to .5 90 +.1 to.2 18

>.5 13 >.2 14

Table 3.2.3.2 Keyrun vs. 5 predators run. Linear models fitting the change in suitability and M2 to the change in predator biomass. Models with both overall slope and with separate slopes for each predator are fit. Where significant fits were found, table entries are percent of variance explained, and, for separate slopes model, the species with significant parameter estimates. Changes are expressed as both absolute value (tonnes) and percent of mean biomass of the two runs.

Change in: suit M2

Predator biomass .007 (+) .011 (+)

Prey biomass NO NO

% Predator biomass .011 (+) .010 (+)

% Pre y B iomass NO NO

Separate slopes:

Predator biomass .239 (- all predators) .031 (+ Whiting

+ Haddock)

Prey biomass .059 (- Haddock) .028 (+ Herring)

%Predator biomass .241 (- all predators) NO

% Prey biomass .071 (- Haddock) .030 (+ Herring

- Norway Pout)

Fixed effects model: .053 .086

Predator species .048 .077

Prey species NO NO

Predator * Prey NO .006

l

Table 3.2.4.1 Frequencies of occurrence of various magnitudes of differences in suitabilities and M2 between KEYRUN and run with horse mackerel added. All predators combined.

MAGNITUDE OF FREQUENCY OF MAGNITUDE OF FREQUENCY OF

CHANGEIN OCCURRENCE CHANGEINM2 OCCURRENCE

SUlT ABILITY

> -0.4 6 >-.4 l

-.2 to 0.4 5 -.2 to -.4 o

-.1 to -.2 6 -.l to -.2 l

-.05 to -.1 18 -.05 to -.1 l

-.01 to -.05 70 -.01 to -.05 3

-.005 to -.01 46 -.005 to -.01 4

-.001 to -.005 119 -.001 to -.005 21

O to -.001 621 O to -.001 383

O to +.001 389 0.0 to +.0001 585

+.001 to .005 50 +.0001 to .001 110

+.005 to .01 24 +.001 to .005 53

+.01 to .05 41 +.005 to .01 33

+.05to.l 9 +.01 to .05 49

+.1 to .2 18 +.05 to .l 20

+.2 to .4 4 +.1 to .2 5

>+.4 5 >+.2 6

Table 3.2.4.2 Keyrun vs. Horse Mackerel run. Linear models fitting the change in suitability and M2 to the change in predator biomass. Models with both overall slope and with separate slopes for each predator are fit. Where significant fits were found, tab le entries are percent of variance explained, and, for separate slopes model, the species with significant parameter estimates. Changes are expressed as both absolute value (tonnes) and percent of mean biomass of the two runs.

Change in: _~u it M2

Predator biomass .058 (+) .030 (+)

Prey biomass .021 (-) NO

% Predator biomass .036 (+) .015 (+)

% Prey Biomass NO NO

Separate slopes:

Predator biomass .201 (Horse Mackerel) .161 (Horse Mackerel)

Prey biomass .067 (Herring) NO

% Predator biomass .206 (Horse Mackerel) .167 (Horse Mackerel)

% Prey biomass .063 (Herring) NO

Fixed effects model: NO .065

Predator species NO .031

Prey species NO NO

Predator * Prey NO .006

E:\ACFM\MAWG97\T-32ll.DOC 10/09/97

Table 3.2.5.1 Frequencies of occurrence of various magnitudes of differences in suitabilities and M2 between KEYRUN and run with old consumption rates. All predators combined.

MAGNITUDE OF FREQUENCY OF MAGNITUDE OF FREQUENCY OF

CHANGEIN OCCURRENCE CHANGEINM2 OCCURRENCE

SUITABILITY

> -0.04 4 >-.l 3

-.02 to -.04 49 -.05 to -.l 8

-.01 to -.02 61 -.Otto -.05 39

-.005 to -.01 86 -.005 to -.01 31

-.001 to -.005 192 -.001 to -.005 110

-.0005 to -.001 70 -.0005 to -.001 54

-.0001 to -.0005 109 -.0001 to -.0005 107

O to -.0001 121 0.0 to -.0001 129

O to +.0001 54 0.0 to +.0001 279

+.0001 to .0005 140 +.0001 to .0005 171

+.0005 to .001 154 +.0005 to .001 72

+.001 to .005 217 +.001 to .005 118

+.005 to .01 63 +.005 to .01 44

+.01 to .02 48 +.01

to

.05 68

>.02 l >.05 14

Table3.2.5.2 Keyrun vs. old consumption run. Linear models fitting the change in suitability and M2 to the change in predator biomass. Models with both overall slope and with separate slopes for each predator are fit. Where significant fits were found, tab le entries are percent of variance explained, and, for separate slopes model, the species with significant parameter estimates. Changes are expressed as both absolute value (tonnes) and percent of mean biomass of the two runs.

Change in: su it M2

Predator biomass .011 (+) .172 (+)

Prey biomass .014 (-) NO

% Predator biomass .011 (+) .132 (+)

% Prey Biomass .052 (-) NO

Separate slopes:

Predator biomass .021 (+ Whiting) .381 (+ Whiting

- Haddock)

Prey biomass .209 (- Haddock NO

- Norway Pout)

c~

% Predator biomass .022 (+ Whiting) .349 (+ Whiting

- Haddock)

% Prey biomass .212 (- Haddock NO

- Whiting - Norway Pout)

Fixed effects model: .018 .034

Predator species NO .029

Prey species .010 NO

Predator * Prey NO NO

E:\ACFM\MAWG97\T-32ll.DOC 10/09/97

Table 4.2.2.1 Sample input data from 'cannibalism' scenario.

Bmax 70Rmax 100

M2 at age maturity oartial weight at ml+ z numbers SSB

l o o 0.05 0.5066 0.5066 100 o

0.5 o O.l O.l 0.3533 0.3733 60.25 o

0.2 o 0.5 0.2 0.2613 0.3613 41.46 o

O.l 0.5 0.8 0.4 0.2307 0.3907 28.90 5.781

o l l 0.6 0.2 0.4 19.56 11.734

o l l 0.8 0.2 0.4 13.11 10.487

o l l l 0.2 0.4 8.79 8.787

o l l 1.2 0.2 0.4 5.89 7.068

o ^{l l 1.4 0.2 0.4 3.95 5.528}

Table 4.2.2.2 Stock recruit parameters for the True and VPA estimated scenarios.

Max Rec. SSB at Max Rec. a b

True 100 70 3.88 .0143

All-age 54 40 3.67 .025

Sequential · 40 61 1.81 .0165

Cannibalism 53 42 3.43 .0238

Figure 2.1.4.1.1

Survey indices of abundance of grey gurnard by size class (in mm) from the IBTS data A. 1st quarter surveys by year

35 - 30

~

25 -

.. :l 20 _._

l ! o ..

& 15 -

z

10 ; 5

~

o

1970 1975 1980 1985

B. Means from quarterly surveys 1991-1995.

140;

! 120 _;_

100 T

.. :l

80 +

l ! o .. CD

Q.

60

z

40 ~

'

20 l

at ^~

2

quarter

1990 1995

3 4

2000

-D-<100 _.._.100-200

-ø- 200-300 -iiE-->300

-D-<100 -.-1oo-200 -&-200-300 ---->300

145

Figure 2.1.4.1.2

Estimated average biomass of grey gurnard by year corresponding to the quarterly input values of MSVPA (dots) based on IBTS data, compared to relative biomass estimates from EGFS (circles).

450000 - 400000- 350000 -

."

300000-

G c

250000

-r

.2 c _{200000 -L}

§ ^{150000 ~}

100000- 50000-

o-

1970 1975 1980 year 1985 1990 1995

Figure 2.1.4.2.1

Survey indices of abundance of starry ra y by size class (in mm) from the IBTS data.

A. 1st quarter surveys by year

3.5- 3 - 2.5

-t-

--o-<100

.. :li 2-

o -la--100-200

.l! .. -&-200-300

CD

a. _{1.5 -} --->300

z

1 - 0.5 _;_

o

1970 1975 1980 1985 1990

B. Means from quarterly surveys 1991-1995.

14 - 12

--o-<100

.. -.-10D-200

:li

o

-&-200-300

.l! ..

--->300

CD

6_;_

z a.

^l

! 4 t 2,

o~ ~ e

2 3 4

quarter

Figure 2.1.4.2.2

Estimated average biomass of stany ray by year corresponding to the quarterly input values of MSVPA (dots) based on IBTS data, compared to relative biomass estimates from EGFS (circles).

300000- 250000 ...

."

200000 ~

c CD

o 150000 - c

- ~ _{100000 ~}

50000 ~

l

o

1970 1975 1980 1985 1990 1995

-1

year

Figure 2.3.1

'OLD' Cod Conaumptton ••Hmat . . 'NEW' Cod Conaumptlon .. tlrnot ..

35 35

30 30

25 25

20 20

:a

o 15

:a

o 15

10 10

5 5

o o

o -

^{N (fl}

""

^It)

^age

^<l

^,.._

^Cl)

^o-

^~

^o -

^{N (fl}

""

^ltQg~

^,.._

^Cl)

^o-

^~

'OLD' WhiHng ConsumpHon esHmates 'NEW' WhiHng ConsumpHon esHmatea

o

o -

^{N (fl}

"" d6e -o ,.._

_{Cl) C>}

~

'OLD' Salthe ConsumpHon estimat., 'NEW' Salthe Conaumptlon estimat ..

25 25

20 20

15 15

:a

o

::t.

o

10 10

5 5

o o

(fl It)

,.._

~e :::

~

!!2

^{(fl It)}

^,.._ _~e

_~

!!2

'OLD' Haddock ConsumpHon .. umatea 'NEW' Haddock Consumptlon .. Hrnotea

Cod PF

-

V\ o Figure 2.4.1 Weight at age.

Cod as predator

12~---~

10

8 l ^• • - ^l _{__.,_Age4}

_,._Age 6

6+ l ~~S4 ^Age7

-+-S6 ....,._S7

:::ir4 ~~

^{L _ _ _}

~~ ^{)( )( )( )(}

^~

~4 ~~4 ^{)( )(} lit ⁾⁽

4 l

2

0+---~---~---~---+---~

1970 1975 1980 1985 1990 1995

Figure 2.4.2

2 1.8 1.6 1.4 1.2 1 0.8 0.6 0.4 0.2

o

1974 1976

V\ -

Hadd PF

Weight at age.

Haddock as predator

1978 1980 1982 1984 1986

Year

Page 1

1988 1990 1992 1994

_.,_Age 4 -Ill-Age 6 Age8

~54

... s6

--4t-S8

Vl N

Mack PF

Figure 2.4.3 W eight at age.

Mackerel as predator

0.7~---

0.6

0.5

+ : : : : : : : : ^{~ ....} 7 ".--=l ^\\. _l

0.41 ~ ~ ~ ~ ~ ~ ~--~ ~ y ^{)( )(} _{~ ~ \><}

^{p • • • ,}

0.3 r

~

0.2

0.1

0+---~---~---r---~---+---~---+---~

1974 1976 1978 1980 1982 1984 1986 1988 1990

Year

-+-Age4 l---Age 6 Age8

~Series4

---seriess

~Series6

Saithe PF

Figure 2.4.4 Weight at age.

Saithe as predator

6 l - l

5

4 l ^{-·- --- --- --- --- --- --- --- --- --- --- --- --- ---}

^{...-.-. .,.}

T

_.,_Age 4 _,._Age6

l Age8

/ --- ",____. --... -

¹

^--*-S4

3+

----ss

-+-S8 2

1

0+---~---~---4---~---~

19 75 1980 1985 1990 1995

Year

-

V\ w

Page 1

VI

~

Figure 2.4.5

0.9 0.8 0.7 0.6 0.5 0.4 0.3 0.2 0.1

o

1974

Weight at age.

1976 1978 1980

Chart2

Whiting as predator

1982 1984 1986 1988

Year

1990 1992 1994

-+-Age 4 -li-Age 6

···=L···.· Age 8 --*-S4 __._S6

---sa

Chart2

Figure 2.4.6 Weight at age. C od

3~---_,

2.5

2 l r

-+-Age 1 --11-age 2

1.5 + ^l

¹

^{· Age3}

-*-Stom 1 ---stom2

~Stom3

l - ^{- ...._ ~ l}

1

0.5

0+---+---+---~---+---~

1970 1975 1980 1985 1990 1995

-

VI VI

Page 1

-

Vl

0\

Figure 2.4. 7 Weight at age.

0.5 0.45 0.4 0.35 0.3 0.25 0.2 0.15 0.1 0.05

o

1974 1976 1978

Chart3

haddock weight at age comparison

1980 1982 1984 1986 1988 1990 1992 1994

---Had 1 Had2

~Had3

--+-Series1

---series5

__._ Series6

Figure 2.4.8

0.2

0.18

0.16

0.14

0.12

C) 0.1

~

0.08

0.06

0.04

0.02

o

1970

- ^VI

-...)

Herring F

W eight at age.

Herring

~ . . . ^~

1975 1980 1985 1990

Year

Page 1

1995

---Age 1 Age2

~Age3

-+-Sto 1

---sto2

-+-Sto 3

Chart1

-

Vl

00

Figure 2.4.9 Weight at age.

Mackerel

0.45

0.4

0.35

0.3 l .____ ^_/ _{\ l}

-+-Age 1

: 0.25 ^0.2 I : : : : : : : : ^{~X )( 4t: )( )( ~ ; ] . _,._Age2 ~Sto1 ---sto2 Age3}

-+-Sto3

l ~ l

0.15

0.1

0.05

o

1974 1976 1978 1980 1982 1984 1986 1988 1990

Year

VI 1.0

NPout F

Figure 2.4.10 Weight at age. N. Pout

60~---~

50

40

l( •.

lK ' .• lK ••... ... ···-.... )K ... /···' ... · J( )K • .

w < ... .v.

J( ·· J( J( ..• .. ... ..:·.-; ... ,.. ~

E 30 æ

20

10

0+---+---r---r---~~---~---~---~

1974 1976 1978 1980 1982 1984 1986 1988

Year

Page 1

---Age 1

···Age2

~Age3

-+-Sto1

-)l-Sto 2

-e-Ste 3

0\ o

æ

Chart4

Figure 2.4.11 W eight at age. Sand Eel

35~---~

30

25

20

15

10

5

0+---+---+---~---~---~---~---~

1974 1976 1978 1980 1982 1984 1986 1988

Year

Plaice Fest

Figure 2.4.12 W eight at age. Plaice est

0.35~---~

0.3

0.25

l ' ^{V ~ ~··· -/\ l} - ^{\./1 -+-Age 1}

0.2-1- lE • • • • • • • • ^-~d ..,.L. _{• • • • • •}

* I-111-Age 2 Age3 --:)E-Sto 1

0.15 l ^{X ~ ~ ~=--. • • • • • • • • • • •} _{~ : :}

=i ^{1--!11-St 2 3}

0.1

0.05

0+---+---+---+---+---4

1970 1975 1980 1985 1990 1995

Year

0\

Page 1

Saithe F

-

0\

Figure 2.4.13 Weight at age.

N

12 ---~

1

0.8

l A ^{r -} _{-+-Age 1}

---Age2

0.6 + - ^{l ~ 1\/ l 1} _~Sto1 ^Age3

_._Sto2 --e-Sto 3

l ~ ~ j ^~ _• - "'l~ _• l \ /l

0.4

0.2

o l---~---~---~---+---~

1970 1975 1980 1985 1990 1995

0'1 w .li:: æ

Chart7

Figure 2.4.14 Weight at age. Sole WEST

o~ ---~

0.2

0.15 + -- ..._ __._ _r '\. /\. ... ^{/ \ ~}

l - ^--.--- • _l

0.1 -l- l

0.05

Or---~---+---r---~---_j

1970 1975 1980 1985 1990 1995

Year

Page 1

1

--+-Age 1 -la-Age2

l ... Age3

~Sto1

__._Sto 2

l-e-sto 3

Chart4

~

Figure 2.4.15 Weight at age.

Weight at age comparison Whiting

0.3~---~

0.25

0.2

0.15

~,.". )l( )l( ··."~·· ,/. )l( ·..•.•. )l( ·....• )l( )l( )l( • )l( '\.' )l( )l( )l( )l( • . . · )l( )l( •'•qq''*

0.1

0.05

0+---~---+---~---~---~

1970 1975 1980 1985 1990 1995

-11- Whiting 1

·· ·q· ^{Whiting 2}

~Whiting3

-+-Sto1

_._Sto2

--e--Sto 3

-

T""

t: CD

cE -o

(.) o

(a6epaJd)s

C'

~--····--··

..

-·---;:~:~F:;·--·----·-·-t-·---

... .

1-- ·l

o ^~-

o·o

(a6epaJd)s

(l ... -J-... ¹

JaiJenb ¹⁶⁵

o

T"""

00

co

Q) Q)

C> O>

ro ro

"O "O

co ~ ~

o. o.

co

T""" T"""

en O>

ri ^-.::t-

t:: ~

Cl)

cE ^~

"O 'O

o o

(.) (.)

o·o ^~'O- g·o z·o-

{a6epaJd)s (a6epaJd)s

~ 1···-"···-···-·l···-·· .. ···1 ~ ... -... -t····---... ..

J ... ,

L

"r'"

O>

&6

~ t::

"'C (.) o

z:·o

(a6epaJd)s

o·o

JeJ,Jenb

<O

OL'O-

'- Q)

~ co

::J O"

""'""

O>

<6 t:: Q)

~ _o

(.)

e·o o·o

(a6epaJd)s

~·o o·o z:·o-

JeJ,Jenb

CX)

(()

167

'- Q)

t:::

(O

::J o-

Q) O) co

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~ c..

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C) c .E

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-··· ...

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T""

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® _o

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L

o·o

(a6epaJd)s

~--···- ^···l

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co _"C ^«J

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J··· ··-···

o

""""

CX)

Q) Cl)

C) O>

co ^«'$

co "' ^~ _o. ^{co ~} _Q.

;•o o·o e·o- o·o

(a5epaJd)s (a6epaJd)s

~-···· ^{_ ....} ,

l 1--1

1-1-1 ^{(\ ~ ...} ... ,

'- '-

~ .@

co ^co

::J ::J

er ^O"

~ , ^{... _.,"J} l ... ... ,

li)

L , ^{... ···}

l 1-

tri N

QJ

""

= s

bl)

g·o o·o ^o·~- v·o o·o ^g·cr

JeJ,Jenb J9l-fenb ¹⁶⁹

-q-

0) c:

'E cu

~ _o u

-.eL

li)

N

~

..

= s

bl)

o·o

(a6epaJd)s

1-· ···l

l··· ···l

l···· ···l

co

L{) O) c:

·c: '-

~ cu _o

(.)

(a6epeJd)s

~-·· ^····l

t··· ·-·1

J-· ····l

co

cu O>

\

co

u co ^~ a.

1···-1-···

" ^{tV ~ ~}

lt)

co

o o

l

- - i - - j_ ...

- ^{(l) en} o ^{o C\1}

~

'- o -

(l) l

!

t:

0-

-

^fl)

cu ^{:::J CTI C\1 o .}

C!

l

j_

. _...

...

l

o co

l

^-l --t- ^;i. - ^{- ..__j}

o 5 10 15

quarter predage

" ^{tV ~ ~} -

. T

C\1 o ^..., --r-

o "!"

- ^Cl) O)

cu ^'-

:

'C C\1 1

!!

Cl)

o ^{t: o}

0-

^l

~

-

^fl)

^{c:r o +} ₁ ^T

o co

..q- o

l

l

-

:6 -

- - _--

2 6 10 14

quarter p red age

'- cu

Cl)

~

'- cu

(l)

~

LO o

l

lt)

...

l

&O

C'li

_l

lt)

o

o

lt) l

... ~

l

co o

l

l

~

..

l

~"

"Q)

~<o

"Q) PJ"

"Q)

:

- -

l

-

r T

: ;

++

_{i ;}

l l

- ^.. --·

year

~"

"qj OJq,<o ~ p_,"-

~ "qj "Qj

-+--

---

"'!"'"

--r-- i

l.

"':"'

!t ~

year

T l

i

--t--

-

i j

-r- .._ T

! .l.

- ^{; .. -} -

year

171

- ^-.l _N

o

"' _c. ^{ål OI}

(Q

m

_~

(1)

o

~

OI

..c c m ::a.

..,

(1)

~ m ..,

s(predage)

-0.6 -0.2 0.2

f- ... _·_ --

^{l l l l}

_l

quarter

-0.4 0.0 0.4

·f-+l

t 1·-··+····1

l+j

~ •·-····-····•···-·i

year

-0.3 -0.1 0.1 0.3

_.________._

• 1--t-1

1·-···1···-·-···-~

1·-·-···l···l

7

~

a

-v

7&

~7 7&

~eS

s(predage) -0.4 -0.2 0.0 4=----~--

^{l l l l l}

o ^..._ ' l

"' ^ål _c. ⁰¹

(Q

m

_~

(1)

o

'

~

-· .-

OI

quarter

-0.5 0.0 0.5 1.0

+

.Q

l···t··l

m c

::a.

₍₁₎

rj-1 ..,

1···+·-~

year

-0.2 0.0 0.2 0.4

• ^1-1·-l

~ ·-- , ^=~i::L• : ::~---+- ---·

~ m ..,

7

..?

a

S'

7&

~7 7&

~eS

"' ^ål _c.

(Q

m

(l)

o

OI

~

o

~

OI

.Q c

IU ::a.

(1)

..,

~ m ..,

s(predage)

-0.6 -0.2 0.2

- ' l

l l l l

l ^(/)

-·-·- --

^'

^{l m} ::s

(1)

c.

9!.

1: '.\'.

t l

\ \

\ \

: '

quarter

-0.2 0.0 0.2

1··+··1 ^{l a}

~ 1--+--jl "

year

-0.2 0.2 0.6

• 1-~l

~ ·--·--·---+-- ··---·

1···~·-····;·.~---··-·+···1

7&

(97 7&

es> eS

(lQ

~

c .,

~

N (.,..

Qc

Figure 3.1.2.1

MS VPA summary for the years ¹⁹⁷⁴ - ¹⁹⁹⁵

Species: C od

North Sea, Key run

500 500

-.. _{. -}

-+--" ₄₀₀ -+--" 400

o o

t:::::)

t:::::)

9 E

- _o::l

( / ) 300 ifJ ₃₀₀

Cl)

( / )

·c;

~ ^Cl)

r:n o..

r:n

^ifJ

ro

^{( / )}

s _o

^{200 ~}

_>-.

²⁰⁰

05 ~~ ....c:>

•

,..!::.::l c::

c.:>

_-+--" Cl)

o

₁₀₀

æs

100

-+--" j:;:c.:l

( / )

-

o o

l l l 1 l l l l l l l l l l 1 1 1 l l l l l 1 1 1 1 l l l 1 l l l 1 1 1 l l l l 1 1 1 1

9 9 9 9 9 g g g g 9 9 9 Q 9 9 g g 9 9 g g 9 g 9 g Q g 9 g 9 g 9 9 Q 9 9 g 9 9 g 9 g g 9

7 7 7 7 7 7 8 8 8 8 8 8 B B B 8 g 9 g g g g 7 7 7 7 7 7 8 8 8 8 8 B B B B 8 g g g g 9 9

4 5 B 7 B g

o

l 2 3 4 5 6 7 8 g o 1 2 3 4 5 4 5 6 7 8 g o 1 2 3 4 5 6 7 8 g o 1 2 3 4

es

bU U bU U

..-...

-+--"

400 ·l

o

⁴⁰⁰

t:::::)

c::::>

..::__

-

^-+--"

_o

³⁰⁰ ifJ ^{Cl) ifJ} ;::J ³⁰⁰

t:::::)

ro

t:::::) C,.)

-::::__...

~

Cl)

'"O ...c:

QJ

200 -+--"

o

200

~ s _o

..._.

~

'"O

100

ro

¹⁰⁰

Cl)

~

-

o o

1 1 1 1 1 1 1 1 1 l l l l l l 1 l l 1 1 l 1 1 1 1 1 1 1 1 l l 1 1 1 1 1 1 1 1 1 1 1 1 l

9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 g 9 9 9 9 9 9 9

7 7 7 7 7 7 8 8 8 a 8 8 B 8 B 8 9 9 9 9 9 9 7 7 7 7 7 7 8 8 8 a 8 B 8 8 B 8 9 9 9 9 9 9

4 (5 6 7 8 9 o 1 2 3 4 5 6 7 B 9 o 1 2 3 4 5 4 (5 6 7 8 9 o 1 2 3 4 5 8 7 8 9 o 1 2 3 4 5

.970 13000

.980 .950

.940 12000

.930 .920

11000 .910

.900

.890 10000

.a8o

.870

..-...

co

.660 ifJ

9000

l

.650

c::

~

C\.1 .840

Q) .830 8000

0.0

^.820

_g

ro

^.810 ₇₀₀₀

!:"=...-

^.800 _.790 ifJ

-+--"

~ ^.780

·s

⁶⁰⁰⁰

ro

^.770 ~

Q) .780 C,.)

~ ^{.750 Cl) ISO OO}

.740

o::

.730

.720 4000

.710 .700

l

3000 .690

.680

l

.670 2000

l l

.660

l l ^l

.6150

l

l

.640 1000 ^l

.

1 1 1 1 1 1 l 1 1 1 l l l l l 1 1 1 1 l l 1 1 1 1 1 1 1 1 1 l l 1 1 l l 1 l l l 1 l l 1

9 g 9 9 9 9 9 g 9 9 g 9 9 9 9 9 9 9 9 g 9 g g g g g g g g g 9 g g g g g 9 g g g g 9 g g

7 7 7 7 7 7 8 B 8 B 8 8 8 B 8 8 g g g g g g 7 7 7 7 7 7 8 8 8 8 8 8 8 B 8 B g g g g g g

4 15 6 7 8 g

o

l 2 3 4 15 6 7 8 g

o

l 2 3 4 5 4 5 6 7 B g o 1 2 3 4 15 6 7 B g o 1 2 3 4 15

Figure 3.1.2.2

MSVPA summary for the years ¹⁹⁷⁴ - ¹⁹⁹⁵

Species: Haddock

North Sea, Key run

l~UU l~OU

lBOO lBOO

1700 1700

_...--.., 1600 1600

~ _...--..,

1500

...,

1500

2:::::> 2:::::>

c::::>

1400

c::::>

¹⁴⁰⁰

c::::>

E

..;:__..

1300 1300

o:::l

¹²⁰⁰

rn

¹²⁰⁰

Cl:)

1100 Cl) 1100

Cl:)

·c;

~ ^{1000 Cl) 1000}

rn

900

P-.

rn rn

900

<:t:l

BOO Cl:) BOD

s ^::::s

o

⁷⁰⁰

>-.

⁷⁰⁰

~

⁶⁰⁰

^...el

^BOD

~ ~

/

500 Cl) 500

<:...>

...,

o

400

<:t:l

⁴⁰⁰

...,

Cl:) ~

300 300

200

l~

²⁰⁰

100 100

o o

l l l

l

l 1 l 1 l 1

l

l l l 1 l 1 l 1

l

l 1 1 1 l 1 l

l

l

l

l 1 1 1

l

1 1 l 1 1 l 1 1 1

g 9 g 9 g g 9 g g g 9 g g g g g g 9 g 9 g 9 g 9 9 9 9 9 9 g 9 g g g g g g g g 9 g 9 g 9

..,

7

..,

7 7

..,

B B B B B B B B B B 9 g g 9 g 9

.., .., .., ..,

7

..,

B a

a

^a a a a a a B g 9 g 9 9 9

4 IS a 7 a g

o

1 2 3 4 IS B

.., _a

g

o

l 2 3 4 IS 4 IS 6 7

a

^g

o l

2 3 4 IS 6

..,

B 9

o

l 2 3 4 IS

J.900· J."'UU

laoo

^{l BOD}

1700 1700

-

1600 ...,:>

laoo

11500

' 2:::::>

¹¹⁵⁰⁰

c::::>

1400

c::::>

¹⁴⁰⁰

..;:__..

1300

U2 ¹³⁰⁰

- ^...,

^{1200 Cl) U2 1200}

2:::::>

¹¹⁰⁰

^;::::1

¹¹⁰⁰

c::::> ^<:t:l

c::::>

¹⁰⁰⁰ <:...> ₁₀₀₀

..;:__..

~

900 Q..) 900

'"d

_BOD

....c::

_BOD

Ci3 ...,

:>=:

⁷⁰⁰

_aoo s o _o

600 ⁷⁰⁰

~ 1500

l~ ^...

¹⁵⁰⁰

400

'"d

₄₀₀

ro

300 ^Q..) 300

~

200 200

100 100

!~

o o -

l

l

1 l l l l 1 l 1 l 1 l l l l l l 1 l l l l

l

l l l l l l l l l l l l 1 l l l l l l

l

9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9

..,

7

..,

7 7

..,

B B B B B B B B B B 9 9 9 9 9 9

..,

7

..,

7 7

..,

B B

a

a a B B a B B 9 9 9 9 9 9

4 IS B 7

a

9

o

1 2 3 4 5 B

..,

B 9

o

l 2 3 4 5 4 IS B 7 B 9

o

l 2 3 4 5 B

..,

B 9

o

1 2 3 4 5

l . l u 11uuuu

1.08

1.06 100000

1.04

1.02 90000

1.00 .9BO

BOOOO

.960

-

co

.940

rn

l

_.920 ~

~

⁷⁰⁰⁰⁰

C\.1 Cl) .900 t:U) .aBo

~

⁶⁰⁰⁰⁰

ro

_.BBO

~- _.640

rn

₆₀₀₀₀

...,:>

~ ^.B20

';:3

<:t:l

_{.800 ~}

Q..) C,) 40000

::21

^{.7BO Cl)}

.'760

o::;

30000 .740

.720

.700 20000

.660

l

.660 10000

l l l ~

Figure 3.1.2.3

MSVPA summary for the years ¹⁹⁷⁴ - ¹⁹⁹⁵

Species: Saithe

North Sea, Key run

>IDD BDO

aoo aoo

.-.. _o ^__,

⁷⁰⁰

- ^..._, _o

⁷⁰⁰

c:::> c:::>

E

600

E

⁶⁰⁰

o:::l en

Lr1 Cl)

Lr1 ~o

o T)

^~o

o

c:!d

^Cl)

en ^0... en

en

_{400 400}

cO

Lr1

s ^::::s

o ::>-.

P5

³⁰⁰

^....0

³⁰⁰

~

Cl

C,.) _{..._..)} Cl)

o

²⁰⁰

_cO

²⁰⁰

__,

~

Lr1

-

100 100

o o

1 l l l 1 1 1 l 1 1 l l l l l 1 l 1 1 1 l 1 l 1 l l l l l l l l l l l 1 1 l 1 1 l l 1 l

9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9

7 7 7 7 7 7

e e a e e a a e e e

9 9 9 9 9 9 7 7 7 7 7 7

e e e e e

B

e e

B

e

9 9 9 9 9 9

4 IS 6 7

e

9

o

1 2 3 4 IS 6 7 B 9

o

l 2 3 4 IS 4 5 6 7 B 9

o

1 2 3 4 IS 6 7 B 9

o

l 2 3 4 IS

'OI OD BOD

BOD BOD

-

^..._..)

700 ·l

o

700

c:::>

E

600 r::n. 600

-

^{..._..) Q.)}

^en

o ^;:::J

c:::>

^~o

^o ro

ISO O

c:::>

^C,.)

~ H

Q.)

""'

⁴⁰⁰

^...c::

⁴⁰⁰

cu ^..._,

!

:;:::: o

300

s _o

³⁰⁰

...

H

200

""' ^ro

^{Q.) 200}

~

~

100 100

o o

l l l l 1 l l l l 1 l l l 1 1 l l 1 l 1 1 1 l l l l 1 1 1 l 1 1 l l l l l 1 1 1 1 1 l 1

9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9

..,

7 7 7 7 7 B B

e e

B B

e

B B

e

9 9 9 9 9 9 7

..,

7 7

..,

7

e

^{B B} B

e

B B B B B 9 9 9 9 9 9

4 ~ 6 7 B 9

o

l 2 3 4 5 6 7 B 9

o

1 2 3 4 IS 4 5 6

.., e

9

o

l 2 3 4 5 6

..,

B 9

o

1 2 3 4 ~

l . DO t:IDO

.900

ISO O

.600

-

\:.O en

l Cl

er:>

Cl) .700

~

⁴⁰⁰

g

0.0 cO

r::...- en

..._..)

Cl

^.600

'9

300

cO

_H

Cl) C,.)

~ ^Cl)

.500 ~

200 .400

l l ^l

.300 _{l l 1 1 1}

l

_{l l}

^l

_{l l l l l l l l 1 l l l l l l} ¹⁰⁰ _l

l

_l

l

_l

^l

_{l 1 l l l l l}

l

_{l l} ^l _{l l l}

l

_l

^l

l l l 1 l 1

g g 9

"

^{9 9 9 9 9 B 9 9 9 9 9 9 9 9 9 9 9 9 9 g 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9}

7 7 7 7 7 7 B B

e

B

e e e

B

e

B 9 9 9 g 9

"

^{7 7 7 7 7 7}

^e

^{B B B B B 8 B 8 B 9 9 9 9 9 9}

4 IS 6 7 B 9

o

l 2 3 4 5 6 7 B g

o

l 2 3 4 IS 4 5 6 7

e

9

o

l 2 3 4

o

6 7 B 9

o

l 2 3 4 ~

Figure 3.1.2.4

MSVPA summary for the years ¹⁹⁷⁴ - 1995

Species: Mackerel

North Sea, Key run

~UUU· ~uuu

900 900

- . . ..., -..

800

...,

₈₀₀

o _o

c:> c:>

s=>

700

E

⁷⁰⁰

- _co

600 1Zl 600

U ) Cl.>

U )

~

T)

Cl.>

!500

0...

^!500

1Zl 1Zl

1Zl

<'lj U )

s

⁴⁰⁰

^:::s

⁴⁰⁰

o :>-.

i::Q

₃₀₀ ^,...Cl 300

,.!::<::1 0 ...,

^~ Cl.>

o

₂₀₀ <'lj 200

...,

U ) ~

100 100

o ~ o

~ 1 l ~ l l 1 1 1 l 1 1 1 1 l 1 l 1 l l l l l l l l 1 l l l 1 l l l l l l l l l l l l l 9 9 Q Q Q Q 9 Q Q Q 9 Q 9 9 9 Q 9 9 9 9 Q 9 9 9 Q 9 9 Q 9 9 9 Q 9 9 9 9 Q 9 Q 9 Q Q Q Q '7' '7' '7' 7 7 '7'

a a a a a a a a a a

^{Q Q} 9 Q Q Q 7 7 '7' '7' '7' '7'

a a a a a a a a a

B Q Q Q Q Q Q 4 IS 6 7 B Q

o

l 2 3 4 IS

e

7

e

^Q

o

l 2 3 4 IS 4 IS 6 7

e

^Q

o

l 2 3 4 IS

e

7

e

9

o

1 2 3 4 IS

luuu· 1000

900 900

- ^...,

aoo

l

o c:>

^BOO

700

~

700

- ^...,

C/) ^{Cl.) C/)}

o

⁶⁰⁰

^;::::!

⁶⁰⁰

c:>

^<'lj

c:> u

..::.._....

ISO O

H

500

Cl.)

""C1

^~

Cl) ...,

400

o

⁴⁰⁰

::::::: s

300

o

300

... H

""C1

200

·~

^{~ <'lj Cl.) 200}

^l~

100 100

o o

l l 1 1 1 l l l l 1 1 l l l 1 ~ l l l ~ l l l 1 l 1 l l 1 l ~ l l 1 l l 1 l ~ l l 1 1 1

9 9 9 Q 9 9 Q 9 9 9 9 9 9 9 9 9 9 Q 9 Q 9 9 9 9 9 9 9 9 9 9 Q 9 9 9 9 9 9 9 9 9 9 9 9 9

7 7 7 7 7 7 B B B B

e a a

B

e

B 9 9 9 9 9 Q '7' 7 '7' 7 7 7 B

a

B

a

^B

a a a a a

9 9 9 9 9 9 4 5 6 7 B 9

o

1 2 3 4 5 6 7

e

9

o

l 2 3 4 5 4 5 6 7 B 9

o

l 2 3 4 5 6 7 B 9

o

l 2 3 4 5

.BOO 600

.700

500

.600

-

c:o

^1Zl ~ 400

l

_.500

~

~ Cl.>

_g

Q.O

<'lj

.400 300

~-

...,

1Zl

~

·a

<'lj

.300

H

Cl.>

0

~ Cl.> 200

o=;

.200

100 .100

1.1111

l l l

Figure 3.1.2.5

MS VPA summary for the years ¹⁹⁷⁴ - ¹⁹⁹⁵

Species: Whiting

North Sea, Key run

1uOu· 1000

900 900

,....-., ^-joooJ

_o

⁸⁰⁰

- _o

^{-joooJ 800}

c:> c:>

c:>

₇₀₀

2:-

⁷⁰⁰

..:::.__...

CCl

600 lf:J ₆₀₀

er:;

Q)

er:; ·c:;

~ ₅₀₀ ^Q)

o..

⁵⁰⁰

lf:J lf:J

en

l

er:;

ro s

^{400 ~ 400}

o ::>-.

i:iS

300

^....a

~ ³⁰⁰

,..!:o::<

_Q)

c..>

-joooJ

o

₂₀₀

ro

200

-joooJ ~

er:;

100 100

o o

l 1 l 1 l 1 l 1 l l 1 l 1 l 1 l 1 l 1 1 1 1 1 1 l 1 1 1 l 1 l 1 1 1 1 l 1 l 1 l l l l 1

9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 g 9 9 9 9 9 g 9

7 7 '7 '7 '7 '7 6 8 8 8 8 8 8 8 8 8 9 9 9 g 9 9 '7 '7 '7 '7 '7 '7 6

a a

B

a

B 8 8 B 8 9 9 9 9 g 9

4 5 6 7 8 9

o

1 2 3 4 (5 6 '7 B 9

o

l 2 3 4 (5 4 5 B '7 8 9

o

1 2 3 4 5 6 7 6 9

o

l 2 3 4 5

l.UUU 1uuu

900 900

-

^-joooJ

800 ·l

o _c:>

⁸⁰⁰

'700

2:-

700

lf:J

-

^{-joooJ Q) lf:J}

o

⁶⁰⁰

^;::::::1

⁶⁰⁰

c:> ro

c:> c..>

..:::.__...

500 h ⁵⁰⁰

Q)

'""d ...c:::

Cl)

400 ^-joooJ

o

⁴⁰⁰

:;::::

l

s

300

o

300

...

h

'""d

200

ro

²⁰⁰

Q)

~

~ ^~

100 100

- -

o o

l 1 l 1 l 1 l 1 1 1 1 l 1 l 1 1 1 l 1 1 l 1 l 1 1 1 1 1 l 1 l 1 1 l 1 1 1 l 1 l 1 1 1 1

9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9

'7 7 '7 7 '7 '7 8 8

a a a

B

a

⁸

a

B 9 9 9 9 9 9 '7 '7 '7 7 7 7 B B B B

a

B B 8 8 B 9 9 9 9 9 9

4 5 6 7 8 9

o

1 2 3 4 5 6 7

a

9

o

1 2 3 4 5 4 5 6 7 8 9

o

1 2 3 4 5 6 '7 8 9

o

l 2 3 4 5

1.;::u BOOOO

1.10

70000

1.00

-

⁶⁰⁰⁰⁰

e.o

^lf:J _~

l

.900

~

02

Q)

_g

⁵⁰⁰⁰⁰

Q.IJ

ro

_.800

~- lf:J

-joooJ

~

·a

⁴⁰⁰⁰⁰

ro

_.700

h

Q) (.)

~ ^Q)

0::

30000

.600

.500 20000

l

.400

l

¹⁰⁰⁰⁰

^{l l} l l

1 l 1 1 1 l l 1 1 1 1 1 1 l 1 1 l 1 1 l 1 1 1 1 l l l 1 1 l 1 1 1 1 1 1 l l 1 1 l l l 1

9 g g g g g 9 g g g g 9 g g g g g 9 g g g 9 g g g g g g g g 9 g g g 9 g 9 g 9 g g 9 g g

7 7 7 7 7 7 8 B 8

a

8 8 8 B B B g 9 g 9 9 g 7 7 7 7 7 7 B B 8 8

a

8 8 B B 8 9 g g g g g

4 5 6 7 8 g

o

1 2 3 4 5 6 7 B g

o

l 2 3 4 (5 4 5 6 7

a

^g

o

l 2 3 4 5 6 7 B 9

o

1 2 3 4 5

Figure 3.1.2.6

MS VPA summary for the years 1974 - 1995

Species: Herring

North Sea, Key run

GUUU c:

o u u

1900 1900

1900 1900

----.

^{1700 1700}

-+->

₁₉₀₀

....-...

-+->

1600

o c:::::>

1500

o c:::::>

1500

9

1400

E

¹⁴⁰⁰

- o::l

_r:/) ¹³⁰⁰ ₁₂₀₀

en

¹³⁰⁰ ₁₂₀₀

<l.>

r:/)

1100

T)

1100

~

<l.>

1000

o..

¹⁰⁰⁰

en en

en

900 900

ro

^r:/)

s

^{800 ~ 800}

o

700

:;:::....,

₇₀₀

p:;

₆₀₀ ^,....c::l

600

~

o

500

<l.>

₅₀₀

C,.)

-+->

o

₄₀₀

ro

₄₀₀

-+->

_~

r:/)

300 300

200 200

100 100

o o

1 l 1 1 1 l 1 1 1 1 1 1 l 1 l 1 1 1 l 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 l 1 1 1 1 l l. l l. l

9 9 9 g 9 9 9 9 9 9 9 9 g g g 9 9 9 9 g 9 9 9 9 9 9 9 9 9 g 9 9 9 9 9 9 9 9 9 9 9 9 9 g

7 7

...,

_{7 7 7 9}

_{a a a}

₆

_{a e}

₈

_a

_{8 9 9 9} g ₉ g 7 7

...,

_{7 7}

...,

₉

_{a a a a}

₈

_a

₈

_a

_{B 9 9 9 9 9} g 4 15

e

7 8 9

o

l 2 3 4 5

e

7

a

9

o

1 2 3 4 5 4 15

e

7

a

9

o l.

2 3 4 5

e

7 8 9

o

1 2 3 4 5

:0:000 2000

1900 1900

l.aoo l.aoo

1700

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