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Philosophiae Doctor (PhD) Thesis 2019: 22

Integrated Weed Management in Teff: Case of Tigray, Ethiopia

Integrert ugrasbekjempelse i kornarten Teff, Tigray, Etiopia

Norwegian University of Life Sciences Faculty of Biosciences

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NorwegianUniversityofLifeSciences,FacultyofBiosciences,DepartmentofPlantSciences(IPVǦNMBU),P.O.Box5003, NOǦ1432Ås,Norway

NorwegianInstituteofBioeconomyResearch(NIBIO),P.O.Box115,NOǦ1431Ås,Norway Ǧƒ‹ŽǣŽƒ”•Ǥ‘Žƒ˜Ǥ„”ƒ†•ƒ‡–‡”̷„—Ǥ‘ȀŽƒ”•Ǥ‘Žƒ˜Ǥ„”ƒ†•ƒ‡–‡”̷‹„‹‘Ǥ‘

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NorwegianInstituteofBioeconomyResearch(NIBIO),P.O.Box115,NOǦ1431Ås,Norway Ǧƒ‹ŽǣŒƒǤ‡–Žƒ†̷‹„‹‘Ǥ‘

”‘ˆ‡••‘”‡•‡”–—‡

NorwegianUniversityofLifeSciences,DepartmentofInternationalEnvironmentandDevelopmentStudies(NoragricǦ NMBU),P.O.Box5003,NOǦ1432Ås,Norway

Ǧƒ‹ŽǣŒ‡•Ǥƒ—‡̷„—Ǥ‘

”‘ˆ‡••‘”އƒ”–‹Ž‘

NorwegianUniversityofLifeSciences,DepartmentofPlantSciences(IPVǦNMBU),P.O.Box5003,NOǦ1432Ås,Norway NorwegianInstituteofBioeconomyResearch(NIBIO),P.O.Box115,NOǦ1431Ås,Norway

Ǧƒ‹Žǣ‘އƒ”–‹Ǥ‡Ž‘̷‹„‹‘Ǥ‘

”Ǥ‹”ƒ›‡•ˆƒ›‡Žƒ›

MekelleUniversity,CollegeofDrylandAgricultureandNaturalResources,P.O.Box231,Mekelle,Tigray,Ethiopia Ǧƒ‹Žǣ‰‹”ƒ›Ǥ–‡•ˆƒ›̷—Ǥ‡†—Ǥ‡–

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••‘…‹ƒ–‡”‘ˆ‡••‘”ƒ‡”–ƒ•–‹ƒƒ•

CropandWeedEcologyGroup,CentreforCropSystemsAnalysis,WageningenUniversity,P.O.Box430,6700AK Wageningen.TheNetherlands.Tel:31Ǧ(0)317Ǧ484770;Fax:31Ǧ(0)317Ǧ484892

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‡‹‘”…‹‡–‹•–”Ǥ—ƒƒŽ‘‡

NaturalResourcesInstituteFinland(Luke),NaturalResourcesUnit,FIǦ31600JOKIOINEN,Finland.Tel.+358295326517

Ǧƒ‹ŽǣŒ—ƒǤ•ƒŽ‘‡̷Ž—‡Ǥˆ‹

‘‹––‡‡…‘‘”†‹ƒ–‘”

”‘ˆ‡••‘”‡ǦŒ‡”•–‹ŠŽ‡

NorwegianUniversityofLifeSciences,FacultyofBiosciences,DepartmentofPlantSciences(IPVǦNMBU),P.O.Box5003, NOǦ1432Ås,Norway

Ǧƒ‹Žǣƒ‡Ǥ—ŠŽ‡̷„—Ǥ‘

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…ƒ’ƒ…‹–›„—‹Ž†‹‰‹…Ž‹ƒ–‡•ƒ”–ƒ‰”‹…—Ž–—”‡‹–Ї‘”‘ˆˆ”‹…ƒdzǤ

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–‡…А‹…ƒŽƒ†’‡”•‘ƒŽƒ†˜‹…‡•ǤƒƒŽ•‘‹†‡„–‡†ƒ†‰”ƒ–‡ˆ—Ž–‘›…‘Ǧ•—’‡”˜‹•‘””Ǥƒ

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†—”‹‰ –Ї ’‡”‹‘† ‘ˆ › •–—†›Ǥ ƒ ‰”ƒ–‡ˆ—Ž –‘ –Šƒ ”‘ˆ‡••‘” އ ƒ”–‹ Ž‘ ˆ‘” Š‹•

˜ƒŽ—ƒ„އ–‡…А‹…ƒŽŠ‡Ž’ƒ†•‘‘‡”ˆ‡‡†„ƒ…†—”‹‰–Ї™”‹–‹‰‘ˆ‘•–‘ˆ–Ї’ƒ’‡”•Ǥƒ

‰”ƒ–‡ˆ—Ž–‘”Ǥ‘”ˆ‹‘”’ˆ‘”Š‹•‰—‹†ƒ…‡ƒ†ˆ‡‡†„ƒ…†—”‹‰†ƒ–ƒƒƒŽ›•‹•ƒ†’ƒ’‡”

™”‹–‹‰Ǥ ‘”–Š‡Žƒ„‘”ƒ–‘”›–‡…А‹…‹ƒ•ƒ–™‡‡†•…‹‡…‡•‡…–‹‘ǡ–Šƒ›‘—˜‡”›—…Š

ˆ‘”›‘—”Їޒ†—”‹‰„‹‘ƒ••ƒ›‡š’‡”‹‡–ǤЇ‰—‹†ƒ…‡†—”‹‰–Ї•–ƒ”–‘ˆ›ˆ‹‡Ž†™‘”

‘™‡‡†‹†‡–‹ˆ‹…ƒ–‹‘ƒ†–Ї…‘‡–•‘›ˆ‹”•–’ƒ’‡”’”‘˜‹†‡†„›‡”‹–—•”‘ˆ‡••‘”

¤”‡”•–‡‹›‡™‡”‡”‡ƒŽŽ›ƒ’’”‡…‹ƒ–‡†ƒ†‹˜ƒŽ—ƒ„އǤŠƒ›‘—’”‘ˆ‡••‘”¤”‡Ǥ

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ƒ––‡”•†—”‹‰›•–ƒ›ƒ–Ǥ–Šƒ†‡”•›•˜‹ǡ‡š’‡”–ƒ–ǡˆ‘”Š‹•

’‡”•‘ƒŽ…‘‹–‡–†—”‹‰’—”…Šƒ•‡‘ˆƒ–‡”‹ƒŽ•”‡“—‹”‡†ˆ‘”›•–—†›ƒ†”‡’ƒ‹”‘ˆ›

Žƒ’–‘’–Š”‘—‰Š‘—–›•–—†›’‡”‹‘†ǤЇ–‡…А‹…ƒŽ•—’’‘”–ˆ”‘‡‹ˆ%‰‡‘ˆ™ƒ•ƒŽ•‘

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”Ǥ ‹”ƒ› ‡•ˆƒ› ‡Žƒ› ™ƒ• •‘ ‹˜ƒŽ—ƒ„އ ƒ† ˜‡”› ‰”ƒ–‡ˆ—ŽǤ – ™‘—ކ ‘– „‡ ‡ƒ•› –‘

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Їƒ”–‹Ž›•—’’‘”–ǤŠƒ›‘—˜‡”›—…Š”Ǥ††‹•„”ƒŠƒǡ”Ǥއ–•‡Šƒ›•‡‰ƒ›ǡ”Ǥ‹”—

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ƒ••‹•–ƒ…‡ ˆ”‘ •–ƒˆˆ• ‘ˆ ‡’ƒ”–‡– ‘ˆ ”›Žƒ† ”‘’ ƒ† ‘”–‹…—Ž–—”ƒŽ …‹‡…‡•ǡ ƒ†

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ˆ‘”–Їˆ‹‡Ž†™‘”‡”•ƒ–š—ƒ†‡‡ŽŽ‡”‡•‡ƒ”…Š•‹–‡•Ǥ

Šƒ˜‡”‡…‡‹˜‡†Š‡ƒ”–ˆ‡Ž–ƒ†˜‹…‡•ǡ‡…‘—”ƒ‰‡‡–ƒ†•Šƒ”‡††‡Ž‹‰Š–ˆ—Ž‘‡–•ˆ”‘ƒ†

™‹–Š›ˆ”‹‡†•‹†ƒ‡ƒ”‹ƒ„”‡ŠƒǡƒŠ•ƒ›‡„”‡ƒ”‹ƒ‹”‡ƒƒ†Š‹•™‹ˆ‡‡™‹ƒ

ƒ†‡ŽŽ‡ǡ‹”ƒ›‡”Їǡ‹”ƒ›ƒŠ•ƒ›ǡƒ™‹–‡„”‡‡œ‰ƒ„Ї”ƒ†Š‹•ˆƒ‹Ž›ǡ•‡†ƒŽ•”‡•ǡ

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‹”ƒ‡•Š „”‡Šƒǡ ƒ‹Ž‡ ‡Ž‡ƒ”‡‰ƒ› ƒ† ƒ™‹– ƒŠ•ƒ› –‘ ‡–‹‘ •‘‡Ǥ Šƒ ›‘—

ˆ”‹‡†•Ǩ

ƒ†‡‡’Ž›ǡЇƒ”–‹Ž›ƒ†’”‘—†Ž›‰”ƒ–‡ˆ—Ž–‘Šƒ˜‡ƒŽ‘˜‡Ž›‘–Ї””‡‰ƒ•Š••‡ˆƒ™Š‘‹•ƒ

•–‡’’‹‰Ǧ•–‘‡ǡ•–”‘‰„ƒ•‡ƒ†ƒ‡›–‘›’”‘‰”‡••‹˜‡•—……‡••‹‡˜‡”›™ƒŽ‘ˆ›Ž‹ˆ‡Ǥ

Šƒ›‘—˜‡”›—…Š›‘ǤƒƒŽ•‘•‘‰”ƒ–‡ˆ—Ž–‘Šƒ˜‡›„”‘–Ї”•ƒ†ƒ––Ї•ƒ‡

–‹‡›…Ž‘•‡ˆ”‹‡†•ǡ‡”Ї”‡ˆƒ›‡ƒ†‡•ƒŽ‡‰„‡„‡ǡ ˆ‘”„‡‹‰…‘”‡”•–‘‡•ƒ†

‘˜‡”ƒŽŽ•–”‡‰–Š‹‡˜‡”›™ƒŽ‘ˆ›Ž‹ˆ‡ǤŠƒ›‘—˜‡”›—…Š‹†‡‡†Ǥ

Aboveall,thankyoumyalmightyGOD!

ƒˆ–ƒ—‡„”‡–•ƒ†‹‡„”‡Š‹™‘–

%•ǡ‘”™ƒ›

ƒ—ƒ”›ǡʹͲͳͻ

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Abstract...viii

Sammendrag...xii

Listofpapers...xvi

1.INTRODUCTION...2

ͳǤͳǤ ƒ…‰”‘—†ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤʹ ͳǤͳǤͳǤ”‘’’”‘†—…–‹‘—†‡”–Š‹‘’‹ƒƒ‰”‹…—Ž–—”‡ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤʹ ͳǤͳǤʹǤЇ…”‘’–‡ˆˆǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶ a. Briefdescription...4

b. Breedingandagronomy...5

c. Itsproductionandconstraints...11

ͳǤͳǤ͵Ǥ‡‡†•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͳ͵ a. Generaldescription...13

b. Impactoncropproductivity...14

c. Control...14

ͳǤͳǤͶǤ–‡‰”ƒ–‡†™‡‡†ƒƒ‰‡‡–ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͳ͹ a. Concept...17

b. Roleincropproductivity...19

c. Limitations...19

ͳǤͳǤͷǤˆŽ—‡…‡‘ˆ…‘˜‡”…”‘’•‘™‡‡†•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤʹͲ ͳǤͳǤ͸ǤŽ‹ƒ–‡…Šƒ‰‡‹–Š‹‘’‹ƒǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤʹͳ a. OverviewandImpact...21

b. Effectonweedmanagement...22

ͳǤʹǤ ”‘„އŒ—•–‹ˆ‹…ƒ–‹‘ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤʹͶ ͳǤ͵Ǥ „Œ‡…–‹˜‡•ƒ†Š›’‘–Ї•‡•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤʹ͸ 2.MATERIALSANDMETHODS...28

ʹǤͳǤ ‡‡†•ƒ’Ž‹‰ǡ‹†‡–‹ˆ‹…ƒ–‹‘ƒ†ƒ’’‹‰ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤʹͺ ʹǤʹǤ ‹‡Ž†‡š’‡”‹‡–•ƒ††ƒ–ƒƒƒŽ›•‹•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤ͵Ͷ ʹǤ͵Ǥ ‹‘ƒ••ƒ›‡š’‡”‹‡–ƒ††ƒ–ƒƒƒŽ›•‹•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤ͵͸ 3.MAINRESULTSANDDISCUSSION...42

͵ǤͳǤ ‡‡†•‹–‡ˆˆˆ‹‡Ž†•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶʹ ͵ǤͳǤͳǤ’‡…‹‡•…‘’‘•‹–‹‘ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶʹ ͵ǤͳǤʹǤ’‘”–ƒ…‡ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶʹ ͵ǤͳǤ͵Ǥ‡’‘”ƒŽƒ†‡…‘Ž‘‰‹…ƒŽ†‹•–”‹„—–‹‘ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶ͵ ͵ǤʹǤ ˆˆ‡…–•‘ˆ–‹ŽŽƒ‰‡ˆ”‡“—‡…›ǡ•‡‡†”ƒ–‡ƒ†‰Ž›’Š‘•ƒ–‡ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶͶ ͵ǤʹǤͳǤ–‡ˆˆ˜‡‰‡–ƒ–‹˜‡ƒ†”‡’”‘†—…–‹˜‡’‡”ˆ‘”ƒ…‡ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶͶ ͵ǤʹǤʹǤ™‡‡††‡•‹–›ǡ†”›™‡‹‰Š–ƒ†…‘˜‡”ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶͷ ͵ǤʹǤ͵Ǥ™‡‡†‹‰–‹‡ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶ͹ ͵Ǥ͵Ǥ ‡‡†…‘’‡–‹–‹˜‡ƒ„‹Ž‹–›‘ˆ–‡ˆˆ˜ƒ”‹‡–‹‡•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶͺ ͵Ǥ͵ǤͳǤ‡‘–›’‹…‡ˆˆ‡…–ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͶͺ ͵Ǥ͵ǤʹǤŽŽ‡Ž‘’ƒ–Š‹…‡ˆˆ‡…–ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͷͳ ͵Ǥ͵Ǥ͵Ǥ’‘”–ƒ–…‘–”‹„—–‹‰–”ƒ‹–•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͷͶ ͵ǤͶǤ ‡‡†•—’’”‡••‹‘ƒ„‹Ž‹–›‘ˆ–‡ˆˆƒ•…‘’ƒ”‡†–‘…‘˜‡”…”‘’•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͷͷ 4.CONCLUSIONANDPRACTICALRECOMMENDATIONS...58

ͶǤͳǤ ‘…Ž—•‹‘ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͷͺ ͶǤʹǤ ”ƒ…–‹…ƒŽ”‡…‘‡†ƒ–‹‘•ǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤǤͷͻ 5.REFERENCE...60

PAPERS...71

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ƒŽ–‹–—†‹ƒŽ”ƒ‰‡•ǡͳͷͲͲǦʹ͵ͲͲ‡–‡”•ƒ„‘˜‡•‡ƒŽ‡˜‡ŽǤ••–ƒ’އ…”‘’–‘‘”‡–ŠƒŠƒŽˆ‘ˆ–Š‹‘’‹ƒ

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Ž‘••ǤЇ‘„Œ‡…–‹˜‡‘ˆ–Š‹••–—†›™ƒ•–™‘ˆ‘ކǡȋ‹Ȍ–‘’”‘˜‹†‡‘™Ž‡†‰‡ƒ„‘—––Ї†‘‹ƒ–™‡‡†•

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Ї •—”˜‡› ™ƒ• …‘†—…–‡† –‘ †‡–‡”‹‡ –Ї ™‡‡† •’‡…‹‡• …‘’‘•‹–‹‘ǡ ‹’‘”–ƒ…‡ ƒ† –Ї‹”

ƒ‰”‘‡…‘Ž‘‰‹…ƒŽ†‹•–”‹„—–‹‘Ǥ‡‡†•’‡…‹‡•…‘’‘•‹–‹‘™ƒ••–—†‹‡†‹ͳʹͺ”ƒ†‘Ž›•ƒ’އ†–‡ˆˆ

ˆ‹‡Ž†•‘ˆ–Їʹ͸ƒŒ‘”–‡ˆˆ‰”‘™‹‰™‡”‡†ƒ•ȋ†‹•–”‹…–•Ȍ‘ˆ‹‰”ƒ›ǤЇˆ‹‡Ž†•™‡”‡•‹–—ƒ–‡†ƒ––Š”‡‡

ƒŽ–‹–—†‡•ǣŽ‘™Žƒ†ȋδͳͷͲͲƒǤ•ǤŽǤȌǡ‹†Žƒ†•ȋͳͷͲͲȂʹͷͲͲƒǤ•ǤŽǤȌƒ†Š‹‰ŠŽƒ†•ȋεʹͷͲͲƒǤ•ǤŽǤȌǤ

ƒ’އ•™‡”‡…‘ŽŽ‡…–‡†ƒ–ƒƒ˜‡”ƒ‰‡‹–‡”˜ƒŽ‘ˆ͵Ȃͷ‹ƒ‹˜‡”–‡†œ‹‰œƒ‰ˆƒ•Š‹‘Ǥ’ƒ–‹ƒŽ

‡š–‡•‹‘‘ˆ™‡‡††‡•‹–›Ž‡˜‡Ž•™‡”‡ƒ’’‡†ƒ†‹–‡”’‘Žƒ–‡†ˆ‘”–Ї‘•–ˆ”‡“—‡–™‡‡†•’‡…‹‡•

–‘ˆ‹‰—”‡‘—––Ї‹”‹’‘”–ƒ…‡‹ƒŽŽ–ЇƒŒ‘”–‡ˆˆ‰”‘™‹‰ƒ”‡ƒ•‘ˆ‹‰”ƒ›—•‹‰Ǥ ”‘ƒŽŽ

–ЇƒŒ‘”–‡ˆˆ‰”‘™‹‰™‡”‡†ƒ•‘ˆ‹‰”ƒ›•—”˜‡›‡†ǡͶʹ™‡‡†•’‡…‹‡•™‡”‡‹†‡–‹ˆ‹‡†ǤŽ–‹–—†‡’Žƒ›‡†

ƒ †‡…‹•‹˜‡ ”‘އ‘—„‡”‘ˆ ™‡‡† •’‡…‹‡•ǤЇЋ‰Š‡•– ƒ˜‡”ƒ‰‡ —„‡”™ƒ•ˆ”‘Š‹‰ŠŽƒ† ȋͳ͹

•’‡…‹‡•Ȁ•ƒ’އȌˆ‘ŽŽ‘™‡†„›‹†Žƒ†ȋͳ͵ǤͶ•’‡…‹‡•Ȁ•ƒ’އȌƒ†Ž‘™Žƒ†ȋͺǤʹ•’‡…‹‡•Ȁ•ƒ’އȌǤЇ

†‘‹ƒ–‹‰ ™‡‡† •’‡…‹‡• ™‡”‡ArgomenemexicanaǡPlantagolanceolate,Cyperusesculentus, Erucastrumabyssinicum,Avenaabyssinica ƒ†GalinsogaparvifloraǤ ˆ –Ї –™‡Ž˜‡ ‘•– ˆ”‡“—‡–

™‡‡†•’‡…‹‡•ǡˆ‘—”™‡”‡’‡”‡‹ƒŽ•ƒ†‡‹‰Š–™‡”‡ƒ—ƒŽ•ǤЇ™‡‡†•’‡…‹‡•™‹–Š™‹†‡•’ƒ–‹ƒŽ

‡š–‡•‹‘ ‘ˆ Š‹‰Š‡” †‡•‹–› އ˜‡Ž• ‹ ‘•– ‘ˆ –Ї ƒŒ‘” –‡ˆˆ ‰”‘™‹‰ ™‡”‡†ƒ• ™‡”‡Erucatrum abyssinicum,Plantagolanceolateƒ†cyperusesculentusǤ

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‡‘ˆ–Їˆ‹‡Ž†‡š’‡”‹‡–•™ƒ•…‘†—…–‡†–‘…Žƒ”‹ˆ›–Ї‡ˆˆ‡…–•‘ˆ–‹ŽŽƒ‰‡ˆ”‡“—‡…›„›ƒ”†

’Ž‘—‰Š‹‰ǡ‰Ž›’Š‘•ƒ–‡ƒ’’Ž‹…ƒ–‹‘’”‹‘”–‘•‘™‹‰–‡ˆˆƒ†•‡‡†”ƒ–‡Ǥ–Š‹•‡š’‡”‹‡–ǡ

–Ї‘•–‹–‡•‹˜‡•‘‹Ž…—Ž–‹˜ƒ–‹‘ǡ…‘’ƒ”‡†™‹–ЖЇއƒ•–‹–‡•‹˜‡ǡ‹…”‡ƒ•‡†‰”ƒ‹›‹‡Ž†

„›ʹ͸Ψǡ–Ї—•‡‘ˆ‰Ž›’Š‘•ƒ–‡‰ƒ˜‡ͳͲ؊‹‰Š‡”‰”ƒ‹›‹‡Ž†ƒ†–Ї–™‘Š‹‰Š‡•–•‡‡†”ƒ–‡•ǡ …‘’ƒ”‡†™‹–ЖЇޑ™‡•–ǡ‹…”‡ƒ•‡†‰”ƒ‹›‹‡Ž†„›ʹ͸ΨǤЇˆ‘ŽŽ‘™‹‰”‡†—…–‹‘•‘ˆ–‘–ƒŽ

™‡‡†„‹‘ƒ••™ƒ•ˆ‘—†Ǣ–Ї‘•–‹–‡•‹˜‡•‘‹Ž–‹ŽŽƒ‰‡ǡ…‘’ƒ”‡†™‹–А‘ƒ”†’Ž‘—‰Š‹‰ǡ

‰ƒ˜‡ƒʹͻΨ”‡†—…–‹‘ǡ–ЇЋ‰Š‡•–•‡‡†”ƒ–‡…‘’ƒ”‡†–‘–Їޑ™‡•–ǡʹͲΨ”‡†—…–‹‘ƒ†

‰Ž›’Š‘•ƒ–‡ •’”ƒ›‹‰ ƒ ͳ͵Ψ ”‡†—…–‹‘Ǥ ‘™‹‰ –‡ˆˆ ‹‡†‹ƒ–‡Ž› ƒˆ–‡” ‘‡ ƒ”† ’Ž‘—‰Š‹‰

ȋ‹‹—–‹ŽŽƒ‰‡Ȍǡƒ–ƒ •‡‡†”ƒ–‡‘ˆ ͳͷ‰ȀŠƒƒ†ƒ’’Ž‹…ƒ–‹‘‘ˆ‰Ž›’Š‘•ƒ–‡ •‹‰‹ˆ‹…ƒ–Ž›

”‡†—…‡† ™‡‡† †‡•‹–›ǡ †”› ™‡‹‰Š– ƒ† …‘˜‡” ™Š‹Ž‡ ‡Šƒ…‹‰ –‡ˆˆ ˜‡‰‡–ƒ–‹˜‡ ƒ†

”‡’”‘†—…–‹˜‡ ’‡”ˆ‘”ƒ…‡Ǥ ”‡“—‡– –‹ŽŽƒ‰‡ ƒ† —•‡ ‘ˆ –Ї ”‡…‘‡†‡† •‡‡† ”ƒ–‡ ȋʹͷ

‰ȀŠƒȌŠƒ†ƒŽ‘•–•‹‹Žƒ””‡•—Ž–•Ǥ

ƒ‘–Ї”ˆ‹‡Ž†‡š’‡”‹‡–ǡ†ƒ–ƒ”‡Žƒ–‡†–‘–‡ˆˆ˜‡‰‡–ƒ–‹˜‡ƒ†”‡’”‘†—…–‹˜‡’‡”ˆ‘”ƒ…‡ǡ

™‡‡††‡•‹–›ǡ„‹‘ƒ••ǡƒ†…‘˜‡”ǡƒ†–‹‡”‡“—‹”‡†ˆ‘”Šƒ†™‡‡†‹‰™‡”‡…‘ŽŽ‡…–‡†ˆ‘”

ͳͲ –‡ˆˆ ˜ƒ”‹‡–‹‡•Ǥ Ї †‹ˆˆ‡”‡– –‡ˆˆ ˜ƒ”‹‡–‹‡• •Š‘™‡† ˜ƒ”‹ƒ–‹‘ ‹ –Ї‹” ’Ї‘Ž‘‰› ȋͷͲΨ

‡‡”‰‡…‡ǡ ˆŽ‘™‡”‹‰ǡ ƒ† ƒ–—”‹–›Ȍǡ Ї‹‰Š–ǡ –‹ŽŽ‡”‹‰ ƒ† ›‹‡Ž†Ǥ ‡–™‡‡ ƒ› ‘ˆ –Ї

˜ƒ”‹‡–‹‡•ǡ–Ї”‡™‡”‡‘‘”‹‘”†‹ˆˆ‡”‡…‡•‹‡ƒ”Ž‹‡••‘ˆ‡‡”‰‡…‡ǡ„—––Ї˜ƒ”‹‡–‹‡•

Ǯ‘•‡–̵ƒ†̵ǦͲͳǦͳ͸ͺͳ̵•Š‘™‡†ǡ‹„‘–Š•‹–‡•ƒ†›‡ƒ”•ǡˆ‡™‡”†ƒ›•–‘ͷͲΨ‡‡”‰‡…‡

–Šƒ̵ǦͲͳǦ͵ͷͶ̵ƒ†̵Ǧ͵ͷͺ̵ǤЇ–™‘˜ƒ”‹‡–‹‡•™‹–Їƒ”Ž‹‡•‡‡”‰‡…‡™‡”‡ƒŽ•‘–Ї

–™‘˜ƒ”‹‡–‹‡•™‹–ЖЇޑ™‡•–’Žƒ–Ї‹‰Š–Ǥ —”–Ї”‘”‡ǡ–Ї–™‘˜ƒ”‹‡–‹‡•™‹–Š–Š‡Žƒ–‡•–

‡‡”‰‡…‡–‹‡ǡ̵ǦͲͳǦ͵ͷͶ̵ƒ†̵Ǧ”Ǧ͵ͷͺ̵ǡ™‡”‡ƒŽ•‘–Ї–™‘˜ƒ”‹‡–‹‡•™‹–ЖЇЋ‰Š‡•–

—„‡” ‘ˆ –‹ŽŽ‡”•Ǥ Ї”‡ ™‡”‡ •‹‰‹ˆ‹…ƒ– †‹ˆˆ‡”‡…‡• ‘ˆ ™‡‡† ‰”‘™–Š ‹ –Ї †‹ˆˆ‡”‡–

˜ƒ”‹‡–‹‡•ǡˆ‘”‡šƒ’އǡ–Ї”‡™‡”‡ͷͷ؎‡••–‘–ƒŽ™‡‡†„‹‘ƒ••‹–Ї–™‘‘•–…‘’‡–‹–‹˜‡

˜ƒ”‹‡–‹‡•ȋ̵ǦͲͳǦʹ͸͹ͷ̵ƒ†̵‘”ƒ̵Ȍ…‘’ƒ”‡†–‘–Ї–™‘އ••…‘’‡–‹–‹˜‡˜ƒ”‹‡–‹‡•ȋ̵ǦͲͳǦ

͵ͷͶ̵ƒ†̵ǦǦ͵ͷͺ̵ȌǤ”ƒ‹›‹‡Ž†‘ˆ–Ї†‹ˆˆ‡”‡–˜ƒ”‹‡–‹‡•˜ƒ”‹‡†…‘•‹†‡”ƒ„Ž›„‡–™‡‡

Ž‘…ƒ–‹‘•ƒ†›‡ƒ”•ǡƒ†‘ˆ–‡‘Ǧ•‹‰‹ˆ‹…ƒ–†‹ˆˆ‡”‡…‡•‘……—””‡†Ǥ̵Ǧ”Ǧ͵ͺ͹̵ƒ†̵‘…ƒŽ̵

™‡”‡ –Ї Š‹‰Š‡•– ›‹‡Ž†‹‰ ˜ƒ”‹‡–‹‡• ™Š‹Ž‡ –Ї –™‘ ˜ƒ”‹‡–‹‡• ™‹–Š –Ї Š‹‰Š‡•– ™‡‡†

‹ˆ‡•–ƒ–‹‘• ȋ̵ǦͲͳǦ͵ͷͶ̵ ƒ† ̵ǦǦ͵ͷͺ̵Ȍ ›‹‡Ž†‡† •‹‰‹ˆ‹…ƒ–Ž› Ž‘™‡” –Šƒ ƒ› ‘ˆ –Ї

‘–Ї” ˜ƒ”‹‡–‹‡• ‹ ʹͲͳͷǤ Ї Š›’‘–Ї•‹• –Šƒ– –‡ˆˆ ˜ƒ”‹‡–‹‡• –Šƒ– ›‹‡Ž† Š‹‰Š ™‹–Š‘—– ™‡‡†

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™ƒ•’ƒ”–‹ƒŽŽ›”‡Œ‡…–‡†„‡…ƒ—•‡–Ї˜ƒ”‹‡–›̵ǦͲͳǦ͵ͺ͹̵‘ƒ˜‡”ƒ‰‡›‹‡Ž†‡†Š‹‰Š‡•–„‘–Š™‹–Š

ƒ†™‹–Š‘—–Šƒ†™‡‡†‹‰Ǥ–Ї‘–Ї”Šƒ†ǡ†‹ˆˆ‡”‡…‡•‹‰”ƒ‹›‹‡Ž†™‹–Šƒ†™‹–Š‘—–

™‡‡†•”ƒ‰‡†ˆ”‘ʹ͹ͳ‰ŠƒǦͳȋ̵‘”ƒ̵Ȍ–‘Ͷ͹ʹ‰ŠƒǦͳȋ̵ǦͲͳǦ͵ͷͶ̵ȌǤ

Š‡Žƒ„‘”ƒ–‘”›‡š’‡”‹‡–™ƒ•†‘‡–‘†‡–‡”‹‡–Ї’‘–‡–‹ƒŽƒŽŽ‡Ž‘’ƒ–Š‹…ƒ…–‹˜‹–›ȋȌ

‘ˆ–‡ˆˆ˜ƒ”‹‡–‹‡••–—†‹‡†‹–Їˆ‹‡Ž†‡š’‡”‹‡–Ǥ–™ƒ•‡ƒ•—”‡†ƒ•–Ї‡ˆˆ‡…–‘ˆ–Ї˜ƒ”‹‡–‹‡•

‘”‘‘–‰”‘™–Š‘ˆ„‘–І‹…‘–ƒ†‘‘…‘–™‡‡†•Ǥ›‡‰”ƒ••ȋLoliumperenne…˜Ǥ‘†‹ƒŽ‡Ǣ …˜α…—Ž–‹˜ƒ”Ȍƒ†”ƒ†‹•ŠȋRaphanussativus…˜ǤЇ””›‡ŽŽ‡Ȍ™‡”‡—•‡†ƒ•‘†‡Ž‘‘…‘–

ƒ† †‹…‘– ™‡‡†• ”‡•’‡…–‹˜‡Ž› ˆ‘” –Ї Žƒ„‘”ƒ–‘”› ȋ„‹‘ƒ••ƒ›Ȍ ‡š’‡”‹‡–Ǥ Ї Žƒ„‘”ƒ–‘”›

‡š’‡”‹‡– •Š‘™‡† –Šƒ– ƒŽŽ –Ї –‡ˆˆ ˜ƒ”‹‡–‹‡• Šƒ† ’‘–‡–‹ƒŽ ƒŽŽ‡Ž‘’ƒ–Š‹… ƒ…–‹˜‹–› ƒ†

‹Š‹„‹–‡†‡ƒ”Ž›”‘‘–‰”‘™–Šƒ††‡˜‡Ž‘’‡–‘ˆ„‘–Š‘‘…‘–ƒ††‹…‘–‘†‡ŽǦ™‡‡†•Ǥ

’ƒ”–‹…—Žƒ”ǡ –Ї Ž‘…ƒŽ ˜ƒ”‹‡–› ȋ̵‘…ƒŽ̵Ȍ „—– ƒŽ•‘ ‘–Ї” ˜ƒ”‹‡–‹‡• •—…Š ƒ• ̵‘•‡–̵ ƒ† ̵ǦͲͳǦ ʹ͸͹ͷ̵™‡”‡ƒ‘‰–Š‘•‡™‹–ЖЇЋ‰Š‡•–ƒŽŽ‡Ž‘’ƒ–Š‹…’‘–‡–‹ƒŽǤ

–Ї•–—†›‘ˆ™Šƒ––”ƒ‹–•–Šƒ–‡š’Žƒ‹–Ї†‹ˆˆ‡”‡…‡‹…‘’‡–‹–‹˜‡‡••„‡–™‡‡˜ƒ”‹‡–‹‡•ǡ

‹–™ƒ•–Ї–Š”‡‡–”ƒ‹–•ƒŽŽ‡Ž‘’ƒ–Š‹…’‘–‡–‹ƒŽǡ–‹‡‘ˆ‡‡”‰‡…‡ƒ†„‹‘ƒ••’”‘†—…–‹‘ǡ

„‡•–‡š’Žƒ‹‡†–Ї†‹ˆˆ‡”‡…‡•„‡–™‡‡–Ї˜ƒ”‹‡–‹‡•ǡ–Š‘—‰ŠŠ‡‹‰Š–‰”‘™–Šƒ†–‹ŽŽ‡”‹‰Šƒ†

ƒŽ•‘˜‹•‹„އ…‘–”‹„—–‹‘Ǥ

‡‡†•—’’”‡••‹˜‡ƒ„‹Ž‹–›‘ˆ–‡ˆˆ™ƒ•…‘’ƒ”‡†™‹–ЖЇ‘•–…‘‘Ž›—•‡†…‘˜‡”…”‘’•

‹ ‹‰”ƒ›Ǥ ‡ˆˆ •—’’”‡••‡† ™‡‡†• ƒ† •‹‰‹ˆ‹…ƒ–Ž› Ž‘™‡”‡† ™‡‡† †‡•‹–›ǡ „‹‘ƒ•• ƒ†

…‘˜‡”ǡ …‘’ƒ”‡† –‘ ˆ‹‡Ž† ’‡ƒ ƒ†Vicia •’Ǥ —…Š ƒ ‡ˆˆ‡…– ™ƒ• …‘•‹•–‡–Ž› ‘„•‡”˜‡† ‹

—™‡‡†‹‰’Ž‘–•Ǥ‘™‡˜‡”ǡ…Šƒ‰‹‰•‘™‹‰‡–Š‘†ˆ”‘„”‘ƒ†…ƒ•–‹‰–‘”‘™•‘™‹‰ƒ†

˜‹…‡˜‡”•ƒ†‹†‘–•‹‰‹ˆ‹…ƒ–Ž›…Šƒ‰‡–‡ˆˆ™‡‡†•—’’”‡••‹˜‡ƒ„‹Ž‹–›Ǥ

‘–Љޑ„ƒŽŽ›ƒ†‹–Š‹‘’‹ƒǡ‹–‹•‹’‘”–ƒ––‘†‡˜‡Ž‘’…”‘’’‹‰’”ƒ…–‹…‡–Šƒ–”‡†—…‡–Ї

‡‡† ˆ‘” ‹–‡•‹˜‡ •‘‹Ž …—Ž–‹˜ƒ–‹‘ ƒ† ˜‘”ƒ…‹‘—• —•‡ ‘ˆ ’‡•–‹…‹†‡•Ǥ  –Š‹• •–—†›ǡ ‹– ™ƒ•

…އƒ”Ž›†‡‘•–”ƒ–‡†–Šƒ–އ••‹–‡•‹˜‡•‘‹Ž–‹ŽŽƒ‰‡”‡•—Ž–‡†‹Š‹‰Š‡”™‡‡†‹ˆ‡•–ƒ–‹‘ƒ†

‰”‡ƒ–‡”‡‡†ˆ‘”ƒ—ƒŽ™‡‡†…‘–”‘ŽŽƒ–‡”‹–Ї‰”‘™‹‰•‡ƒ•‘Ǥ’”ƒ›‹‰™‹–Љޛ’Š‘•ƒ–‡

„‡ˆ‘”‡•‘™‹‰‘ˆ–‡ˆˆ”‡†—…‡†™‡‡†‰”‘™–Š„—–‘––‘–Ї•ƒ‡‡š–‡–ƒ•–Ї‘•–‹–‡•‹˜‡

•‘‹Ž–‹ŽŽƒ‰‡Ǥ‘™‡˜‡”ǡ–Ї•–—†›•Š‘™‡†–Šƒ––Ї…‘’‡–‹–‹˜‡‡••‘ˆ–‡ˆˆ…ƒ•‹‰‹ˆ‹…ƒ–Ž›„‡

‹’”‘˜‡†–Š”‘—‰Š–Ї—•‡‘ˆ…‘’‡–‹–‹˜‡˜ƒ”‹‡–‹‡•ƒ†Š‹‰Š•‡‡†”ƒ–‡•Ǥ‡…‘…Ž—†‡–Šƒ–

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„‘–Š ˜ƒ”‹‡–› •‡Ž‡…–‹‘ ƒ† Š‹‰Š‡” •‡‡† ”ƒ–‡• ƒ”‡ ‹’‘”–ƒ– –‘‘Ž• ˆ‘” ‹–‡‰”ƒ–‡† ™‡‡†

ƒƒ‰‡‡–ƒ†”‡†—…‡–Ї‡‡†ˆ‘”‹–‡•‹˜‡•‘‹Ž…—Ž–‹˜ƒ–‹‘Ǥ

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6DPPHQGUDJ

‡ˆˆȋEragrostistefȋ—……ǤȌ”‘––‡”Ȍ‡”‡•¤ˆ”Þ‡–‘”ƒ”–•‘Š‘˜‡†•ƒ‡Ž‹‰„Ž‹”†›”‡–‹

–‹‘’‹ƒǤ ¤†‡ ‘”ƒ˜Ž‹‰‡ ‡ ‘‰•¤ ŠƒŽƒ˜Ž‹‰‡ ‡” ˜‹–‹‰Ǥ ‡ˆˆ ‡” ‡ ‡––¤”‹‰ǡ •‡Ž˜Ǧ

’‘ŽŽ‹‡”–ǡ ͶǦ’Žƒ–‡ •‘ –‹ŽŠÞ”‡” ‰”ƒ•ˆƒ‹Ž‹‡ ȋPoaceaeȌǤ ‡ –”‹˜‡• „‡•– —†‡” ”‡Žƒ–‹˜–

˜ƒ”‡ˆ‘”Бކ‘‰†‡–‡•–‡ƒ˜’”‘†—•Œ‘‡‹–‹‘’‹ƒˆ‘”‡‰¤”‹‘”¤†‡”•‘Ž‹‰‰‡”ͳͷͲͲǦ ʹ͵ͲͲ‡–‡”‘˜‡”Šƒ˜‡–Ǥ‡ˆˆ—–‰ŒÞ”‡˜‹–‹‰‘‰ƒ˜‰ŒÞ”‡†‡†‡Žƒ˜‘•–Бކ‡––‹Ž‡”‡

ŠƒŽ˜’ƒ”–‡ ƒ˜ „‡ˆ‘Ž‹‰‡ ‹ –‹‘’‹ƒ ‘‰ ‹ ŠÞ›‡”‡Ž‹‰‰‡†‡ ‘”¤†‡” ƒ˜ ”‹–”‡ƒǤ

––‡”•’Þ”•‡Ž‡ ލ‡” „¤†‡ ‹‡Žƒ†• ‘‰ ‰Ž‘„ƒŽ–ǡ ‹‡ ‹•– ˆ‘”†‹ –‡ˆˆ ‹ ‘•–Бކ‡– „Ž‹”

„‡–”ƒ–‡– •‘ Їޕ‡‡••‹‰ ‰—•–‹‰ǡ „Žƒ– ƒ‡– ˆ‘”†‹ †‡ ‡” —–‡ ‰Ž—–‡Ǥ ‡––‡ Šƒ”

ˆ‘”¤”•ƒ‡– ƒ– „¤†‡ ’”‘†—•Œ‘•ƒ”‡ƒŽ ‘‰ Ǧ˜‘Ž— ލ‡” ˆ”ƒ ¤” –‹Ž ¤”Ǥ  ƒ˜ †‡ •–‘”‡

ˆŽƒ•‡ŠƒŽ•‡‡‹’”‘†—•Œ‘‡‡”—‰”ƒ•ǡ„¤†‡ˆ‘”†‹—‰”ƒ•‡–‘—””‡”‘‰†‡”‡†”‡†—•‡”‡”

ƒ˜Ž‹‰‡‡‡‘‰•¤ˆ‘”†‹†‡–ˆ‘”¤”•ƒ‡”›‡ƒ”„‡‹†•”‡˜‡†‡ƒ—‡ŽŽ„‡Œ‡’‡Ž•‡Ǥ‡––‡

‰‹”ˆÞމ‡Ž‹‰„ސ†‡‡ŠÞ›‡ƒ”„‡‹†•”ƒˆ–‘•–ƒ†‡”‘‰Žƒ˜’”‘†—–‹˜‹–‡–Ǥ

¤Ž•‡–‹‰‡‡††‡––‡ŠǦ•–—†‹‡–Šƒ”˜§”––‘Ǧ†‡Ž–ǡȋ‹Ȍ¤ˆ”‡•ƒˆˆ‡—•ƒ’‘Š˜‹Ž‡

—‰”ƒ•ƒ”–‡”•‘‡”†‘‹‡”‡†‡‹–‡ˆˆǦ’”‘†—•Œ‘‡‘‰ȋ‹‹Ȍ¤ˆ‹‡‡–‘†‡”ˆ‘”‹–‡‰”‡”–

—‰”ƒ•„‡Œ‡’‡Ž•‡ ‹ –‡ˆˆ •Ž‹ ƒ– ‡ ƒ Ї˜‡ „§”‡”ƒˆ–‹‰ ’”‘†—–‹˜‹–‡– ‘‰ ”‡†—•‡”‡

’”‘†—•Œ‘•‘•–ƒ†‡‡Ǥ ‘” ¤ ¤ †‹••‡ ¤Ž•‡–‹‰‡‡ Šƒ” †‡– „Ž‹–– ‰Œ‡‘ˆÞ”– „¤†‡

ƒ”–އ‰‰‹‰••–—†‹‡”‘‰ˆ‡Ž–ˆ‘”•ލ‹‘”¤†‡”Š˜‘”–‡ˆˆ†›”‡•‹‹‰”ƒ›ǡ–‹‘’‹ƒ‹ʹͲͳͷ‘‰ʹͲͳ͸Ǥ

ŽŽ‡ˆ‡Ž–ˆ‘”•ލ‡‡„Ž‡Žƒ‰–‘’’•‘•’Ž‹–Ǧ’Ž‘–ˆ‘”•ލ‡†–”‡„Ž‘‡”Ž‘ƒŽ‹•‡”––‹Žš—‘‰

‡‡ŽŽ‡Ǥƒ„‘”ƒ–‘”‹‡ˆ‘”•ލ‡‡„އ—–ˆÞ”–’¤ƒ’—•%•ȋȀȌǤ

‘” ¤ ˆ”‡•ƒˆˆ‡ —•ƒ’ ‘ Š˜‹Ž‡ —‰”ƒ•ƒ”–‡” •‘ ‡” ˜ƒŽ‹‰ ¤ ˆ‹‡ ‹ –‡ˆˆ „އ †‡–

‰Œ‡‘ˆÞ”– ‡ ƒ”–އ‰‰‹‰ ƒ˜ •ƒ‡•‡–‹‰ ‘‰ —–„”‡†‡Ž•‡ ƒ˜ —‰”ƒ•ƒ”–‡” ‹ –‡ˆˆ ‹ —Ž‹‡

ŠÞ›†‡•‘‡”‹‹‰”ƒ›Ǥ‡––‡„އ—–ˆÞ”–’¤–‹Ž•ƒ‡ͳʹͺ–‹Žˆ‡Ž†‹‰’”Þ˜‡”—–‡”ˆ‘”†‡Ž–’¤ʹ͸

ƒ˜ †‡ ˜‹–‹‰•–‡ †‹•–”‹–‡‡ ˆ‘” –‡ˆˆǦ’”‘†—•Œ‘Ǥ ”Þ˜‡”—–‡‡ „އ ˆ‘”†‡Ž– ‹‡ˆ‘” –”‡

ŠÞ›†‡•‘‡”ǣȋͳȌδͳͷͲͲ‘ŠǤǡȋʹȌͳͷͲͲȂʹͷͲͲ‘ŠǤ‘‰ȋ͵ȌεʹͷͲͲ‘ŠǤ”Þ˜‡”—–‡‡„އ

Žƒ‰–‡†‹–‡”˜ƒŽŽ’¤…ƒǤ͵Ǧͷ‹‡–Ǯ•‹•ƒǯÞ•–‡”Ǥ–„”‡†‡Ž•‡‘‰–‡––Ї–ˆ‘”†‡—Ž‹‡

—‰”ƒ•ƒ”–‡‡ „އ ”‡‰‹•–”‡”– ‘‰ ˆ”ƒ•–‹Ž– ’¤ ƒ”– ˜‡† ŠŒ‡Ž’ ƒ˜ †ƒ–ƒ’”‘‰”ƒ‡– ˆ‘”

—Ž‹‡ †‹•–”‹–‡‡Ǥ ¤ †‡ —†‡”•ލ–‡ ‘”¤†‡‡ „އ Ͷʹ —‰”ƒ•ƒ”–‡” ‹†‡–‹ˆ‹•‡”–Ǥ ޛ†‡•‘‡

(15)

•’‹Ž–‡ ‡ ƒ˜‰ŒÞ”‡†‡ ”‘ŽŽ‡ ’¤ ƒ–ƒŽŽ —‰”ƒ•ƒ”–‡”Ǥ ޛ†‡•‘‡ ε ʹͷͲͲ ‘ŠǤ Šƒ††‡

‰Œ‡‘•‹––Ž‹‰ͳ͹ƒ”–‡”ǡ‡––‡”ˆ—މ–ƒ˜•‘‡ʹ‡†ͳ͵ǡͶƒ”–‡”‘‰•‘‡ͳδͳͷͲͲ‘ŠǤŠƒ††‡

ͺǡʹƒ”–‡”Ǥ‡‡•–†‘‹‡”‡†‡—‰”ƒ•ƒ”–‡‡˜ƒ”Argomenemexicana,Plantagolanceolata, Cyperusesculentus,Erucastrumabyssinicum,AvenaabyssinicaogGalinsogaparvifloraǤ˜†‡

–‘Ž˜˜ƒŽ‹‰•–‡—‰”ƒ•ƒ”–‡‡˜ƒ”ˆ‹”‡ˆŽ‡”¤”‹‰‡‘‰¤––‡‡––¤”‹‰‡Ǥ‰”ƒ•ƒ”–‡‡‡†•–‘””‘Ž‹‰

—–„”‡†‡Ž•‡ ‘‰ ŠÞ› –‡––Ї– ˜ƒ”Erucatrumabyssinicum,PlantagolanceolataogCyperus esculentus.

–– ƒ˜ ˆ‡Ž–ˆ‘”•ލ‡‡„އ —–ˆÞ”–ˆ‘” ¤ •‡ ’¤‡ˆˆ‡–‡ ƒ˜ Œ‘”†ƒ”„‡‹†‹‰•ˆ”‡˜‡•‡ ‡† ƒ”†

’Ž‘‰ǡ•’”Þ›–‹‰‡†‰Ž›ˆ‘•ƒ–ˆÞ”•¤‹‰‘‰•¤ˆ”ޏ‡‰†‡ƒ˜–‡ˆˆǤ†‡––‡ˆ‘”•ލ‡–‰ƒ†‡‡•–

‹–‡•‹˜‡Œ‘”†ƒ”„‡‹†‹‰ƒǡ•ƒ‡Ž‹‰‡–‡†‹•–‹–‡•‹˜‡ǡލ–‘”—–„›––‡‡†ʹ͸ΨǤ

’”Þ›–‹‰ ‡† ‰Ž›ˆ‘•ƒ– ‰ƒ ͳͲ Ψ ŠÞ›‡”‡ ‘”—–„›––‡ ‘‰ †‡ –‘ ŠÞ›‡•–‡ •¤ˆ”ޏ‡‰†‡‡

•ƒ‡Ž‹‰‡–‡††‡Žƒ˜‡•–‡Þ–‡‘”—–„›––‡–‡†ʹ͸ΨǤ ގ‰‡†‡”‡†—•Œ‘‡”ƒ˜†‡

–‘–ƒŽ‡—‰”ƒ•„‹‘ƒ••‡„އˆ—‡–Ǣ‡‡•–‹–‡•‹˜‡Œ‘”†„‡ƒ”„‡‹†‹‰‡•ƒ‡Ž‹‰‡–‡†

‹‰‡ ƒ”† ’ŽÞ›‹‰ǡ ‰ƒ˜ ‡ ”‡†—•Œ‘ ’¤ ʹͻ Ψ ƒ˜ —‰”ƒ•‡–Ǥ ‡ ŠÞ›‡•–‡ •¤ˆ”ޏ‡‰†‡

•ƒ‡Ž‹‰‡–‡††‡Žƒ˜‡•–‡‰ƒ˜ʹͲΨ”‡†—•Œ‘ǡ‡•‰Ž›ˆ‘•ƒ–•’”Þ›–‹‰ˆÞ”•¤‹‰‰ƒ˜

‡”‡†—•Œ‘’¤ͳ͵Ψƒ˜—‰”ƒ•‡–Ǥ¤‹‰ƒ˜–‡ˆˆ—‹††‡Ž„ƒ”–‡––‡”‡ƒ”†’ŽÞ›‹‰ȋ‹‹—

Œ‘”†„‡ƒ”„‡‹†‹‰Ȍǡ ‡ •¤ˆ”ޏ‡‰†‡ ’¤ ͳͷ ‰ Ȁ Šƒ ‘‰ „”— ƒ˜ ‰Ž›ˆ‘•ƒ– ˆÞ” •¤‹‰ ‰ƒ˜ ‡

•‹‰‹ˆ‹ƒ– ”‡†—•Œ‘ ƒ˜ —‰”ƒ•–‡––Ї–ǡ –Þ””˜‡– ‘‰ †‡‹‰ǡ •ƒ–‹†‹‰ •‘ †‡– ˆ‘”„‡†”‡–

ƒ˜Ž‹‰‡ƒ˜–‡ˆˆ„¤†‡˜‡‰‡–ƒ–‹˜˜‡•–‘‰ˆ”Þ’”‘†—•Œ‘Ǥ›’’‹‰Œ‘”†„‡ƒ”„‡‹†‹‰‘‰„”—ƒ˜

ƒ„‡ˆƒŽ–•¤ˆ”ޏ‡‰†‡’¤ʹͷ‰ȀŠƒŠƒ††‡‡•–‡Ž‹‡‰‘†‡”‡•—Ž–ƒ–‡”Ǥ

‡– ƒ‡– ˆ‡Ž–ˆ‘”•ލ „އ ͳͲ •‘”–‡” ƒ˜ –‡ˆˆ —†‡”•ލ– ‡† Ї•› ’¤ ˜‡‰‡–ƒ–‹˜ ˜‡•– ‘‰

ˆ”Þƒ˜Ž‹‰ǡ •ƒ– –‹ŽŠÞ”‡†‡ —‰”ƒ•–‡––Ї–ǡ „‹‘ƒ••‡ǡ †‡‹‰ ‘‰ –‹† •‘ ”‡˜‡• ˆ‘”

Ф†Ž—‹‰Ǥ ‡ ˆ‘”•Œ‡ŽŽ‹‰‡ –‡ˆˆ•‘”–‡‡ ˜‹•–‡ ˜ƒ”‹ƒ•Œ‘ ‹ †‡”‡• ˆ‡‘Ž‘‰‹ ȋͷͲΨ •’‹”‹‰ǡ

„Ž‘•–”‹‰ǡ‘†‹‰ȌǡŠÞ›†‡ǡ„—•‹‰‘‰ƒ˜Ž‹‰Ǥ‡ŽŽ‘ƒ‰‡ƒ˜•‘”–‡‡˜ƒ”†‡–‹‰‡

‡ŽŽ‡”‹†”‡ˆ‘”•Œ‡ŽŽ‡”‹–‹†Ž‹‰Š‡–ƒ˜•’‹”‹‰ǡ‡•‘”–‡‡̵‘•‡–̵‘‰̵ǦͲͳǦͳ͸ͺͳ̵Šƒ††‡ˆ‘”

„‡‰‰‡Ž‘ƒŽ‹–‡–‡‡‘‰ˆ‘”•Œ‡ŽŽ‹‰¤”ǡˆ§””‡†ƒ‰‡”–‹ŽͷͲΨ•’‹”‹‰‡̵ǦͲͳǦ͵ͷͶ̵‘‰̵Ǧ

͵ͷͺ̵Ǥ ‡ –‘ •‘”–‡‡ ‡† –‹†Ž‹‰•– •’‹”‹‰ ˜ƒ” ‘‰•¤ †‡ –‘ •‘”–‡‡ ‡† Žƒ˜‡•– ’Žƒ–‡ŠÞ›†‡Ǥ

‹†‡”‡˜ƒ”†‡–‘•‘”–‡‡•‘•’‹”–‡•‡‹‡•–ǡ̵ǦͲͳǦ͵ͷͶ̵‘‰̵Ǧ”Ǧ͵ͷͺ̵ǡ‘‰•¤†‡–‘•‘”–‡‡

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•‘”–‡‡Ǥ ‘”‡•‡’‡Ž˜ƒ”†‡–ͷͷ؏‹†”‡—‰”ƒ•„‹‘ƒ••‡‹†‡–‘‡•–‘—””ƒ•‡•–‡”‡

•‘”–‡‡ ̵ǦͲͳǦʹ͸͹ͷ̵ ‘‰ ̵‘”ƒ̵ •ƒ‡Ž‹‰‡– ‡† †‡ –‘ ‹†”‡ ‘—””ƒ•‡•–‡”‡

•‘”–‡‡ ̵ǦͲͳǦ͵ͷͶ̵ ‘‰ ̵ǦǦ͵ͷͺ̵Ǥ ”Þƒ˜Ž‹‰‡ ƒ˜ †‡ ˆ‘”•Œ‡ŽŽ‹‰‡ •‘”–‡‡ ˜ƒ”‹‡”–‡

„‡–›†‡Ž‹‰‡ŽŽ‘•–‡†‡”‘‰¤”ǡ‡‘ˆ–‡˜ƒ”†‡–‹‡Ǧ•‹‰‹ˆ‹ƒ–‡ˆ‘”•Œ‡ŽŽ‡”Ǥ‹†Ž‡”–‹†˜ƒ”

̵Ǧ”Ǧ͵ͺ͹̵‘‰̵‘…ƒŽ̵†‡‡†ŠÞ›‡•–ƒ˜Ž‹‰ǡ‡•†‡–‘•‘”–‡‡‡†‡•–—‰”ƒ•˜ƒ”̵Ǧ ͲͳǦ͵ͷͶ̵‘‰̵ǦǦ͵ͷͺ̵‘‰‰ƒ˜•‹‰‹ˆ‹ƒ–Žƒ˜‡”‡ˆ”Þƒ˜Ž‹‰‡ƒ‰‡ƒ˜†‡ƒ†”‡•‘”–‡‡

‹ ʹͲͳͷǤ ›’‘–‡•‡ ‘ ƒ– –‡ˆˆ•‘”–‡‡ •‘ ‰‹” •–‘” ‘”ƒ˜Ž‹‰ —–‡ —‰”ƒ•‘—””ƒ•‡ ‰‹”

Ž‹–‡ƒ˜Ž‹‰¤”†‡„Ž‹”—–•ƒ––ˆ‘”—‰”ƒ•’¤‰”—ƒ˜†¤”Ž‹‰‘—””ƒ•‡‡˜‡„އ†‡Ž˜‹•ƒ˜˜‹•–

†ƒ•‘”–‡̵ǦͲͳǦ͵ͺ͹̵‹‰Œ‡‘•‹––‰ƒ˜ŠÞ›‡•–‘”ƒ˜Ž‹‰„¤†‡‡†‘‰—–‡Š¤†Ž—‹‰Ǥ

¤†‡ƒ‡•‹†‡˜ƒ”‹‡”–‡ˆ‘”•Œ‡ŽŽ‡”‹‘”ƒ˜Ž‹‰‡†‘‰—–‡—‰”ƒ•ˆ”ƒʹ͹ͳ‰ŠƒǦͳˆ‘”

̵‘”ƒ̵–‹ŽͶ͹ʹ‰ŠƒǦͳˆ‘”̵ǦͲͳǦ͵ͷͶ̵Ǥ

ƒ„‘”ƒ–‘”‹‡ˆ‘”•ލ‡– „އ —–ˆÞ”– ˆ‘” ¤ „‡•–‡‡ ’‘–‡•‹‡ŽŽ ƒŽŽ‡Ž‘’ƒ–‹• ƒ–‹˜‹–‡– ȋȌ ˆ”ƒ

–‡ˆˆ•‘”–‡‡•‘„އ„”—–‹ˆ‡Ž–ˆ‘”•ލ‡‡‹–‹‘’‹ƒ‘‰‡ˆˆ‡–‡”’¤”‘–—–˜‹Ž‹‰‡Š‘•„¤†‡

–‘ˆ”Þ„Žƒ†‡–‡‘‰‡ˆ”Þ„Žƒ†‡–‡—‰”ƒ•Ǥƒ‹‰”ƒ•ȋLoliumperenne…˜Ǥ‘†‹ƒŽ‡Ǣ…˜α…—Ž–‹˜ƒ”Ȍ‡

‡ˆ”Þ„Žƒ†‡–ƒ”–‘‰”‡††‹ȋRaphanussativus…˜ǤЇ””›‡ŽŽ‡Ȍ‡–‘ˆ”Þ„Žƒ†‡–ƒ”–„އ„”—–

•‘‘†‡ŽŽ—‰”ƒ•Ǥƒ„‘”ƒ–‘”‹‡ˆ‘”•ލ‡–˜‹•–‡ƒ–ƒŽŽ‡†‡–‡•–‡†‡–‡ˆˆ•‘”–‡‡Šƒ††‡ƒŽŽ‡Ž‘’ƒ–‹•

’‘–‡•‹ƒŽ‡‘‰ƒ–†‡Š‡‡––‹†Ž‹‰”‘–˜‡•–‘‰—–˜‹Ž‹‰Š‘•„¤†‡†‡–‡––Ǧ‘‰–‘Ǧˆ”Þ„Žƒ†‡–

‘†‡ŽǦ—‰”ƒ•‡–Ǥ’‡•‹‡Ž–†‡Ž‘ƒŽ‡•‘”–‡Ǯ‘…ƒŽǯ‡‘‰•¤ƒ†”‡•‘”–‡”•‘Ǯ‘•‡–ǯ‘‰ǮǦ ͲͳǦʹ͸͹ͷǯ˜ƒ”„Žƒ–†‡‡†ŠÞ›‡•–ƒŽŽ‡Ž‘’ƒ–‹•’‘–‡•‹ƒŽ‡Ǥ•–—†‹‡ƒ˜Š˜‹Ž‡‡‰‡•ƒ’‡”

Š‘• –‡ˆˆ•‘”–‡” •‘ ˆ‘”Žƒ”‡” ˆ‘”•Œ‡ŽŽ ‹ ‘—””ƒ•‡‡˜‡ ˜ƒ” †‡– †‡ –”‡ ‡‰‡•ƒ’‡‡

ƒŽŽ‡Ž‘’ƒ–‹• ’‘–‡•‹ƒŽ‡ǡ ‘’’•’‹”‹‰•–‹†•’—– ‘‰ „‹‘ƒ••‡’”‘†—•Œ‘ •‘ „‡•– ˆ‘”Žƒ”–‡

ˆ‘”•Œ‡ŽŽ‡”‡ŽŽ‘•‘”–‡‡ǡ‡ŠÞ›†‡˜‡•–‘‰„—•‹‰•‡˜‡„‹†”‘‘‰•¤Ǥ

‘ ‡ †‡Ž ƒ˜ †‘–‘”‰”ƒ†••–—†‹‡– „އ †‡– ‘‰•¤ —–ˆÞ”– ˆ‘”•ލ Š˜‘” †›”‹‰ ƒ˜ –‡ˆˆ „އ

•ƒ‡Ž‹‰‡– ‡† ƒ†”‡ —Ž–—”˜‡•–‡” ˆ‘” ˆ‘”•Œ‡ŽŽ‡” ’¤ —‰”ƒ•ˆ”‡˜‡•–‡Ǥ ‹••‡

ˆ‘”•ލ‡‡‹‰¤”‹‡‹†‡ˆ‹”‡ƒ—•”‹’–‡‡‡‘‡”‡•—Ž–ƒ–‡”‡”˜‹•–‹ƒ˜Šƒ†Ž‹‰‡•

‹Ž‡†‹‰Ǥ ˜‡ –‹Ž –‡ˆˆ ¤ Š‹†”‡ —‰”ƒ•˜‡•– „އ •ƒ‡Ž‹‰‡– ‡† †‡ ‡•– „”—–‡

†‡˜‡•–‡‡‹‹‰”ƒ›Ǥ‡ˆˆ”‡†—•‡”–‡•‹‰‹ˆ‹ƒ–„‡†”‡—‰”ƒ•–‡––Ї–ǡ„‹‘ƒ••‡‘‰†‡‹‰ǡ

•ƒ‡Ž‹‰‡–‡†ˆ‹‡Ž†’‡ƒ•‘‰Viccia••’Ǥ•Ž‹‡ˆˆ‡–„އ•›•–‡ƒ–‹•‘„•‡”˜‡”–‹”—–‡”

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•‘‹‡„އޗ‡–Ǥ†”‹‰ƒ˜ •¤‡–‘†‡ǡˆ”ƒ„”‡‹•¤‹‰ –‹Ž”ƒ†•¤‹‰ǡ‡†”‡–‹‡ –‡ˆˆ•‹

‘—””ƒ•‡‡˜‡Ǥ

¤†‡‰Ž‘„ƒŽ–‘‰‹–‹‘’‹ƒ‡”†‡–˜‹–‹‰¤ˆ‹‡†›”‹‰•‡–‘†‡”•‘”‡†—•‡”‡”„‡Š‘˜‡–ˆ‘”

Œ‘”†ƒ”„‡‹†‹‰•ƒ–‹†‹‰•‘„‡Š‘˜‡–ˆ‘”„”—ƒ˜’Žƒ–‡˜‡”‹†Ž‡”‹‹‡”‡•Ǥ‡—˜‡–‡–

•¤˜‹‹†‡––‡•–—†‹‡–ƒ–‹†”‡‹–‡•‹˜Œ‘”†ƒ”„‡‹†‹‰‘‰•¤‰ƒ‡”—‰”ƒ•‘‰•–Þ””‡„‡Š‘˜ˆ‘”

ƒ—‡ŽŽ—‰”ƒ•„‡Œ‡’‡Ž•‡•‡‡”‡‹˜‡•–•‡•‘‰‡Ǥ’”Þ›–‹‰‡†‰Ž›ˆ‘•ƒ–ˆÞ”•¤‹‰ƒ˜–‡ˆˆ

”‡†—•‡”–‡—‰”ƒ•‡‰†‡‡‹‡‹•ƒ‡‰”ƒ†•‘†‡‡•–‹–‡•‹˜‡Œ‘”†ƒ”„‡‹†‹‰‡Ǥ

–—†‹‡–˜‹•‡”‹‹†Ž‡”–‹†ƒ–‘—””ƒ•‡‡˜‡–‹Ž–‡ˆˆƒˆ‘”„‡†”‡•„‡–›†‡Ž‹‰‰Œ‡‘„”—

ƒ˜‘—””ƒ•‡•–‡”‡•‘”–‡”‘‰•–Þ””‡•¤ˆ”ޏ‡‰†‡”Ǥ‹‘Ž—†‡”‡”ƒ–„¤†‡•‘”–•˜ƒŽ‰‘‰

•¤ˆ”ޏ‡‰†‡‡”˜‹–‹‰‡˜‡”–Þ›„¤†‡ˆ‘”‹–‡‰”‡”–„‡Œ‡’‡Ž•‡ƒ˜—‰”ƒ•‘‰ˆ‘”¤”‡†—•‡”‡

„‡Š‘˜‡–ˆ‘”Œ‘”†ƒ”„‡‹†‹‰Ǥ

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/LVWRISDSHUV

PaperI:

. ƒˆ–ƒ—,J.B.Aune,J.Netland,O.M.EkloandL.O.Brandsæter.’‡…‹‡•

…‘’‘•‹–‹‘ǡ‡…‘Ž‘‰‹…ƒŽ†‹•–”‹„—–‹‘ƒ†‹’‘”–ƒ…‡‘ˆ™‡‡†•‹–‡ˆˆˆ‹‡Ž†•‘ˆ‹‰”ƒ›ǡ

‘”–Ї”–Š‹‘’‹ƒǤƒ—•…”‹’–Ǥ

PaperII:

G.Haftamu,J.B.Aune,O.M.Eklo,T.TorpandL.O.Brandsæter.ˆˆ‡…– ‘ˆ–‹ŽŽƒ‰‡

ˆ”‡“—‡…›ǡ•‡‡†”ƒ–‡ƒ†‰Ž›’Š‘•ƒ–‡ƒ’’Ž‹…ƒ–‹‘‘–‡ˆˆƒ†™‡‡†•‹‹‰”ƒ›ǡ–Š‹‘’‹ƒǤ

ƒ—•…”‹’–Ǥ

PaperIII:

G.Haftamu,J.B.Aune,J.Netland,T.Torp,O.M.Eklo,andL.O.Brandsæter.‡‡†

…‘’‡–‹–‹˜‡ƒ„‹Ž‹–›‘ˆ–‡ˆˆ˜ƒ”‹‡–‹‡•Ǥƒ—•…”‹’–Ǥ

PaperIV:

G.Haftamu,J.B.Aune,O.M.Eklo,T.Torp,andL.O.Brandsæter.ŽŽ‡Ž‘’ƒ–Š‹…’‘–‡–‹ƒŽ

‘ˆ–‡ˆˆ˜ƒ”‹‡–‹‡•ƒ†–Ї‹”‡ˆˆ‡…–‘™‡‡†‰”‘™–ŠǤƒ—•…”‹’–Ǥ

(19)

,1752'8&7,21

%DFNJURXQG

&URSSURGXFWLRQXQGHU(WKLRSLDQDJULFXOWXUH

–Š‹‘’‹ƒ †‡’‡†• ‘ ƒ‰”ƒ”‹ƒ ‡…‘‘› ƒ† ’”ƒ…–‹…‡ ƒ‰”‹…—Ž–—”‡ •‹…‡ ‹–• ‡š‹•–‡…‡Ǥ

‰”‹…—Ž–—”‡…‘–”‹„—–‡•–‘‘”‡–ŠƒŠƒŽˆ‘ˆ‹–•‰”‘••†‘‡•–‹…’”‘†—…–ȋȌƒ†‡’Ž‘›•

‘”‡–ŠƒͺͲΨ‘ˆ‹–•’‘’—Žƒ–‹‘ȋʹͲͲ͹ǢʹͲͳ͸ȌǤЇ…‘—–”›ǡ™‹–Š‹–•Žƒ†ƒ”‡ƒ‘ˆ

ͳǡͳʹ͹ǡͳʹ͹•“—ƒ”‡‹Ž‘‡–”‡•ȋͳͳʹǤ͵‹ŽŽ‹‘Š‡…–ƒ”‡•Ȍǡ—•‡†–‘†‡’‡†‘–Ї•‡…–‘”ˆ‘”ͷͷ Ψ‘ˆ‹–•ȋ‡–ƒŠ—ͳͻ͹ͺȌǤ‘™‡˜‡”ǡƒ‰”‹…—Ž–—”‡‹•…—””‡–Ž›…‘–”‹„—–‹‰͵ͷǤͺΨ‘ˆ–Ї

ȋ™‘”ކˆƒ…–„‘‘ʹͲͳͺȌǤ–Š‹‘’‹ƒƒ‰”‹…—Ž–—”‡…‘•–‹–—–‡•–™‘•›•–‡•ȋͳȌ‹š‡†

…”‘’ȂŽ‹˜‡•–‘…ƒ‰”‹…—Ž–—”‡‹–ЇЋ‰ŠŽƒ†ƒ†Ž‘™Žƒ†ǡƒ†ȋʹȌ’ƒ•–‘”ƒŽ•›•–‡‘•–Ž›‹

–Їޑ™Žƒ†’ƒ”–‘ˆ–Ї…‘—–”›ȋ‡–ƒŠ—ͳͻ͹ͺǢʹͲͳ͸ȌǤЇ‹š‡†…”‘’’‹‰•›•–‡‹•

…Šƒ”ƒ…–‡”‹œ‡†„›Š‹‰ŠŽƒ„‘—”‹’—–ǡŽ‘™’”‘†—…–‹˜‹–›ƒ†—•‡‘ˆ–”ƒ†‹–‹‘ƒŽˆƒ”‹’އ‡–•

ƒ† ƒ‰”‘‘‹… ’”ƒ…–‹…‡•Ǥ ‘” ˆ‘‘† ƒ† ˆ‡‡† •—’’Ž›ǡ ’”‘†—…–‹‘ ‘ˆ …”‘’• ‹• ”‘—–‹‡Ž›

’”ƒ…–‹…‡†‹–™‘•‡ƒ•‘•ƒ—ƒŽŽ›Ǥ•‘‡‘ˆ–Ї…‘’‘‡–•‘ˆƒ‰”‹…—Ž–—”‡ǡ…”‘’’”‘†—…–‹‘

‹•–Ї•‘—”…‡‘ˆ”ƒ™ƒ–‡”‹ƒŽ•ˆ‘”ƒ‰”‹…—Ž–—”ƒŽ‹†—•–”‹‡•ǤЇ…‘—–”›‘™•™‹†‡”ƒ‰‡‘ˆ

ƒ‰”‘Ǧ…Ž‹ƒ–‹……‘†‹–‹‘•ˆƒ˜‘”ƒ„އˆ‘”‰”‘™‹‰–”‘’‹…ƒŽǡ•—„–”‘’‹…ƒŽƒ†–‡’‡”ƒ–‡…”‘’•Ǥ–

‹•ƒŽ•‘–Ї…‡–‡”‘ˆ†‹˜‡”•‹–›ˆ‘”ƒ›…”‘’•ƒ†Š‘•–•Žƒ”‰‡—„‡”‘ˆŽ‹˜‡•–‘…Ǥ

”‘’• ƒ”‡ ’”‘†—…‡† ‹ „‘–Š ”ƒ‹Ǧˆ‡† ƒ† ‹””‹‰ƒ–‹‘ ‡˜‡”› ›‡ƒ”Ǥ ……‘”†‹‰ –‘ …‡–”ƒŽ

•–ƒ–‹•–‹…ƒŽ ƒ—–Š‘”‹–› ȋȌǡ –Ї …‘—–”› Šƒ• ƒ„‘—– ͳ͸ ‹ŽŽ‹‘ Ї…–ƒ”‡• ‘ˆ ƒ”ƒ„އ Žƒ†•

•—‹–ƒ„އ ˆ‘” –Ї ’”‘†—…–‹‘ ‘ˆ ƒ—ƒŽ ƒ† ’‡”‡‹ƒŽ …”‘’• ȋ ʹͲͳ͸Ȍ ™‹–Š ’‘–‡–‹ƒŽ –‘

”‡ƒ…Š—’–‘ͷͷ‹ŽŽ‹‘Š‡…–ƒ”‡•ȋƒ‘„‡‡–ƒŽǤʹͲͲ͹ȌǤˆ–Ї•‡ƒ”ƒ„Ž‡Žƒ†•ǡ‘”‡–Šƒ

ͳʹǤͷ ‹ŽŽ‹‘ Ї…–ƒ”‡• ȋ͹ͺǤͳΨȌ ƒ”‡ ˆ‘” ”ƒ‹Ǧˆ‡† …”‘’ ’”‘†—…–‹‘ ƒ† ͳǤͺ ‹ŽŽ‹‘ Ї…–ƒ”‡•

ȋͳͳǤ͵ΨȌƒ”‡ˆ‘”‹””‹‰ƒ–‹‘ƒ†‘ˆˆǦ•‡ƒ•‘…”‘’’”‘†—…–‹‘ȋʹͲͳ͸ȌǤ”‘’•’”‘†—…‡†‘

”ƒ‹Ǧˆ‡† ƒ† ‹””‹‰ƒ–‡† ƒ”ƒ„އ Žƒ†• ƒ”‡ …Žƒ••‹ˆ‹‡† ƒ• ‰”ƒ‹ …”‘’• ȋ…‡”‡ƒŽ•ǡ ’—Ž•‡•ǡ ƒ† ‘‹Ž

…”‘’•Ȍǡ˜‡‰‡–ƒ„އ•ƒ†”‘‘–…”‘’•Ǥ”ƒ‹…”‘’•ƒ”‡–Ї‘•–‹’‘”–ƒ–ˆ‹‡Ž†…”‘’•ǡ‘……—’›‹‰

ƒ”‘—† ͺ͸ Ψ ‘ˆ –Ї ƒ”ƒ„އ Žƒ† ƒ”‡ƒ ȋ ʹͲͳ͸Ȍ ƒ† ƒ”‡ ƒ‹ •–ƒ› ‘ˆ ‘•– –Š‹‘’‹ƒ•Ǥ

‡”‡ƒŽ• ‘……—’› ͳͳǤ͵ ‹ŽŽ‹‘ Ї…–ƒ”‡• ȋ͹ͻ Ψ ‘ˆ –Ї –‘–ƒŽ ƒ”ƒ„އ Žƒ†•Ȍǡ ’—Ž•‡• ʹǤͳ ‹ŽŽ‹‘

Ї…–ƒ”‡•ȋͳͶǤ͹ΨȌǡƒ†‘‹Ž…”‘’•ͲǤͻ‹ŽŽ‹‘Š‡…–ƒ”‡•ȋ͸Ǥ͵ΨȌǤЇ’”‹…‹’ƒŽ‰”ƒ‹…”‘’•ƒ”‡

–‡ˆˆǡ™Š‡ƒ–ǡ„ƒ”އ›ǡ…‘”ǡ•‘”‰Š—ǡƒ†‹ŽŽ‡–Ǥ‡ˆˆ‰”‘™•‹‘•–ƒ‰”‘‡…‘Ž‘‰‹…ƒŽœ‘‡•‘ˆ

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–Ї…‘—–”›Ǥ‘™‡˜‡”ǡ˜‡‰‡–ƒ„އ•ƒ†”‘‘–…”‘’•ƒ”‡’”‘†—…‡††—”‹‰–Ї‘ˆˆǦ•‡ƒ•‘ƒ‹Ž›

—•‹‰‹””‹‰ƒ–‹‘ƒ†…‘˜‡”͵ǤͲʹΨƒ†ͳ͵ǤͳʹΨǡ”‡•’‡…–‹˜‡Ž›ǡ‘ˆ–Їƒ”ƒ„Ž‡Žƒ†•Ǥ

”‘’’”‘†—…–‹‘—†‡”–Ї…‘˜‡–‹‘ƒŽƒ‰”‹…—Ž–—”‡‹–Š‹‘’‹ƒ‹•„‡‰ƒ„›‹–‡•‹˜‡ƒ†

ˆ”‡“—‡––‹ŽŽƒ‰‡—•‹‰–Ї–”ƒ†‹–‹‘ƒŽardplough(knownas‘mahreshaorקס״؇‹‹‰”‹‰ƒ

ƒ†Ǯmaresha’orק״؇inAmharic)™‹–Šƒ‘„Œ‡…–‹˜‡–‘…‘–”‘Ž™‡‡†•ƒ†…Šƒ”ƒ…–‡”‹œ‡†„›

Ž‘™‹’—–—•‡ǡŽ‘™’”‘†—…–‹˜‹–›ƒ†Š‹‰Š•‘‹ŽŽ‘••„›”—‘ˆˆǤŠ‹•Šƒ•ǡ…‘•‡“—‡–Ž›ǡ”‡•—Ž–‡†

‹•‘‹Žƒ†‡˜‹”‘‡–ƒŽ†‡‰”ƒ†ƒ–‹‘ȋ”ƒ›ƒ‡–ƒŽǤʹͲͳͳǢ”ƒ›ƒ‡–ƒŽǤʹͲͳʹǢƒ—†”‘‡–ƒŽǤ ʹͲͳͶȌƒ†ƒ‡…”‘’’”‘†—…–‹‘˜—ސ‡”ƒ„އ–‘…Ž‹ƒ–‡…Šƒ‰‡ȋЇ‘†‘”‡–ƒŽǤʹͲͳʹȌǤЋއ

•—•–ƒ‹‹‰ƒ‰”‹…—Ž–—”ƒŽ’”‘†—…–‹˜‹–›ƒ†ƒ‹–ƒ‹‹‰•‘‹Ž”‡•‘—”…‡•–Š”‘—‰Š•‘‹Ž—Ž…Š‹‰

ƒ†”‡†—…–‹‘‘ˆ”—‘ˆˆǡ …‘•‡”˜ƒ–‹‘ƒ‰”‹…—Ž–—”‡ȋȌ‹•…‘•‹†‡”‡†ƒ• ƒƒŽ–‡”ƒ–‹˜‡–‘

”‡…Žƒ‹•‘‹Ž†‡‰”ƒ†ƒ–‹‘ƒ†ƒ†ƒ’–…Ž‹ƒ–‡…Šƒ‰‡‹–Š‹‘’‹ƒǤ‹…Ž—†‡•‹‹—•‘‹Ž

†‹•–—”„ƒ…‡ǡ‹…”‡ƒ•‹‰‘”‰ƒ‹…•‘‹Ž—Ž…Šƒ†…”‘’†‹˜‡”•‹ˆ‹…ƒ–‹‘ƒŽ‘‰™‹–ЖЇ‘’–‹—

ƒ’’Ž‹…ƒ–‹‘‘ˆƒ‰”‘‘‹…’”ƒ…–‹…‡•ȋ‘—‰Š‡‘—”ʹͲͲͻǢƒ••ƒ‡–ƒŽǤʹͲͲͻǢЇ‘†‘”‡–ƒŽǤ ʹͲͳʹǢƒ—†”‘‡–ƒŽǤʹͲͳͶǢ”‘—†‡”Ƭ‘‡œǦƒ…’Ї”•‘ʹͲͳͶǢ ƒ”‘‘“Ƭ‹††‹“—‡ʹͲͳͷȌǤ

‹•‰Ž‘„ƒŽŽ›’”ƒ…–‹…‡†‹‘”‡–Šƒͳʹͷ‹ŽŽ‹‘Š‡…–ƒ”‡•‹ƒŽŽ…‘–‹‡–•ȋЇ‘†‘”‡–ƒŽǤ ʹͲͳʹȌǤ•—„ǦƒŠƒ”ƒˆ”‹…ƒ‹…Ž—†‹‰–Š‹‘’‹ƒǡ–Ї”‡‹•Ž‘™ƒ†‘’–‹‘‘ˆ†—‡–‘Š‹‰Š

‹…‹†‡…‡ ‘ˆ ™‡‡†• ȋ‘…•–”ڏ ‡– ƒŽǤ ʹͲͲͻǢ ›••‡ ‡– ƒŽǤ ʹͲͳͳǢ ƒŽ„—‡ƒ ‡– ƒŽǤ ʹͲͳʹȌǤ

Ž‘„ƒŽŽ›ǡ™‡‡†•ƒ”‡–Ї„‘––އ‡…‹ȋƒŽ‡™ƒŒƒʹͲͲ͵Ǣ‘…•–”ڏ‡–ƒŽǤʹͲͲͻǢ ƒ”‘‘“‡–

ƒŽǤʹͲͳͳǢŠƒ—Šƒ‡–ƒŽǤʹͲͳʹǢ”‹…Šƒ”†‡–ƒŽǤʹͲͳ͵Ǣ”‘—†‡”Ƭ‘‡œǦƒ…’Ї”•‘ʹͲͳͶǢ

•Žƒ‹ǡǤǤʹͲͳͶǢ–‡˜‡•‘‡–ƒŽǤʹͲͳͶǢ‹…Бޕ‡–ƒŽǤʹͲͳͷǢ‹‰Š‡–ƒŽǤʹͲͳͷȌǤ‘™‡˜‡”ǡ

Šƒ•’‘•‹–‹˜‡‡ˆˆ‡…–‘„‘–Š•‘‹ŽŠ‡ƒŽ–Šƒ†…”‘’’”‘†—…–‹˜‹–›‹–Š‹‘’‹ƒƒŽ‘‰™‹–Šƒ

‡ˆˆ‹…‹‡– …‘–”‘Ž ‘ˆ ™‡‡†• ȋ‘…•–”ڏ ‡– ƒŽǤ ʹͲͲͻǢ ”ƒ›ƒ‡– ƒŽǤ ʹͲͳͳǢ ›••‡ ‡–ƒŽǤ ʹͲͳͳǢ

ƒ—†”‘ ‡– ƒŽǤ ʹͲͳͶȌǤ Ž‹‡ –‘ ‘–Ї” …”‘’•ǡ –‡ˆˆ Šƒ† Ž‘™‡” ›‹‡Ž† ™‹–Š •‘‡ ‘ˆ –Ї

’”ƒ…–‹…‡•ƒ•‹–™ƒ•˜‡”›•‡•‹–‹˜‡–‘™‡‡†•‡‡”‰‡†‹”‡•’‘•‡–‘–Ї”‡†—…‡†–‹ŽŽƒ‰‡‘

‡”–‹•‘Ž•‹‹‰”ƒ›ȋ”ƒ›ƒ‡–ƒŽǤʹͲͳͳȌǤ

Ї„‹‰‰‡•–…ŠƒŽŽ‡‰‡–‘…”‘’’”‘†—…–‹‘‹–Š‹‘’‹ƒ–‘†ƒ›ƒ”‡„‹‘Ž‘‰‹…ƒŽ–Š”‡ƒ–•ȋ™‡‡†•ǡ

‹•‡…– ’‡•–• ƒ† †‹•‡ƒ•‡•Ȍǡ —•‡ ‘ˆ –”ƒ†‹–‹‘ƒŽ ˆƒ” ‹’އ‡–•ǡ އ•• —•‡ ‘ˆ ƒ‰”‹…—Ž–—”ƒŽ

–‡…А‘Ž‘‰‹‡•ǡ Ž‘™ ‹’—– —•‡ǡ ƒ† ’‘’—Žƒ–‹‘ ’”‡••—”‡ ȋ‡–ƒŠ— ͳͻ͹ͺǢ †‡‡™ ʹͲͲͶǢ

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ƒ‘„‡‡–ƒŽǤʹͲͲ͹Ǣ‡”‡••ƒƬƒ••ƒʹͲͲͻǢ‡‡•‰‡Ƭƒ•Š‹†ʹͲͲͻǢ›••‡‡–ƒŽǤʹͲͳͳǢ

އƒ›‘Š ‡– ƒŽǤ ʹͲͳʹǢ ƒŽ„—‡ƒ ‡– ƒŽǤ ʹͲͳʹǢ ƒ—†”‘ ‡– ƒŽǤ ʹͲͳͶǢ ƒ••‹‡ ‡– ƒŽǤ ʹͲͳͶǢ

–‡˜‡•‘‡–ƒŽǤʹͲͳͶȌǤŠ‹•„‡…‘‡•‘”‡…‘’Ž‹…ƒ–‡†™Š‡…‘„‹‡†™‹–ЖЇ‡‰ƒ–‹˜‡

‡ˆˆ‡…–• ‘ˆ …Ž‹ƒ–‡ …Šƒ‰‡ •—…Š ƒ• •Š‘”–ƒ‰‡ ƒ† —‡˜‡Ž› †‹•–”‹„—–‡† ”ƒ‹ˆƒŽŽǡ ”‡…—””‡–

†”‘—‰Š–ƒ†Ž‘‰†”›•’‡ŽŽ•ȋ‡”‡••ƒƬƒ••ƒʹͲͲͻǢƒ••‹‡‡–ƒŽǤʹͲͳͶȌǤ–‡‰”ƒ–‡†™‡‡†

ƒƒ‰‡‡–‹”‡•’‘•‡–‘–Ї’”‡˜ƒ‹Ž‹‰…Ž‹ƒ–‡…Šƒ‰‡‹•ƒ•‘Ž—–‹‘–‘–Ї•‡…ŠƒŽŽ‡‰‡•

ƒ• ‹– ‹…Ž—†‡• ‹’”‘˜‡† ƒ‰”‘‘‹… ’”ƒ…–‹…‡• ƒ† …”‘’ ˜ƒ”‹‡–‹‡• –Šƒ– …ƒ ‡Šƒ…‡ …”‘’

’”‘†—…–‹˜‹–›ˆ”‘ƒ—‹–ƒ”ƒ„Ž‡Žƒ†Ǥ 7KHFURSWHII a. Brief description

‡ˆˆ ȋEragrostistef(—……)”‘––‡”)‹• ƒ‘‰ –Ї •ƒŽŽǦ‰”ƒ‹‡† …‡”‡ƒŽ …”‘’• †‘‹ƒ–Ž›

‰”‘™‹–Š‹‘’‹ƒˆ‘”‹–•‰”ƒ‹ƒ†•–”ƒ™Ǥ–„‡Ž‘‰•–‘–Їˆƒ‹Ž›‘ƒ…‡ƒ‡…‘•‹•–‹‰‘ˆ͵ͷͲ

•’‡…‹‡•ˆ”‘–Ї‰‡—•”ƒ‰”‘•–‹•ȋ‘•–ƒœƒͳͻ͹ͶǢ‘•–ƒœƒ‡–ƒŽǤͳͻ͹ͻǢ‡–‡ƒͳͻͻ͹ȌǤЇ

‰‡—•‡”ƒ‰”‘•–‹•„‡Ž‘‰•–‘–Ї–”‹„‡Eragrosteae,•—„ˆƒ‹Ž›Eragrostoidae‘ˆ–Їˆƒ‹Ž›

Poaceaeȋ‘•–ƒœƒͳͻ͹ͶȌǤЇ…”‘’Šƒ•ƒ–‹›•‡‡†•™‹–Š•‹œ‡ͳȂͳǤ͹Ž‘‰ƒ†ͲǤ͸Ȃͳ

‹ †‹ƒ‡–‡” ȋƒ„–‡‰‡„”‹ƒŽ ‡– ƒŽǤ ʹͲͲ͹„Ǣ ƒ›‡ ʹͲͳͶȌǤ –Š‹‘’‹ƒ ‹• –Ї …‡–”‡ ‘ˆ ‘”‹‰‹ ƒ†

†‹˜‡”•‹–›‘ˆ–‡ˆˆȋƒ˜‹Ž‘˜ͳͻͷͳȌǤ–‹•„‡Ž‹‡˜‡†–‘„‡‘”‹‰‹ƒ–‡†‹„‡–™‡‡ͶͲͲͲƒ†ͳͲͲͲ

™‹–ЖЇˆ‹”•–•‡‡†•†‹•…‘˜‡”‡†‹ƒ’›”ƒ‹†‹͵͵ͷͻȋƒ˜‹Ž‘˜ͳͻͷͳǢ‘•–ƒœƒͳͻ͹ͶȌǤ ”‘ƒŽŽ‘ˆ–Ї‰‡—•”ƒ‰”‘•–‹•ǡͶ͵Ψ‘”‹‰‹ƒ–‡†‹ˆ”‹…ƒǡͳͺΨ‹‘—–Š‡”‹…ƒǡͳʹΨ‹

•‹ƒǡͳͲΨ‹—•–”ƒŽ‹ƒǡͻΨ‹‡–”ƒŽ‡”‹…ƒǡ͸Ψ‹‘”–Š‡”‹…ƒƒ†ʹΨ‹—”‘’‡

ȋ‘•–ƒœƒͳͻ͹ͶȌǤ‡ˆˆ‹•ƒ•‡ŽˆǦ’‘ŽŽ‹ƒ–‡†ǡƒ—ƒŽǡ™ƒ”•‡ƒ•‘Ͷ‰”ƒ••’”‘†—…‡†ˆ‘”‹–•

ˆ‘‘†ƒ†ˆ‡‡†ȋ‹”Šƒ—‡–ƒŽǤʹͲͳͺȌǤ–‹•ƒ†ƒ’–‡†–‘™‹†‡ƒ‰”‘‡…‘Ž‘‰‹…ƒŽ•‡––‹‰•™‹–Š„‡•–

‰”‘™–Š’‡”ˆ‘”ƒ…‡‹–Ї‹†ƒŽ–‹–—†‹ƒŽ”ƒ‰‡•‹Ǥ‡ǤͳͷͲͲǦʹ͵ͲͲ‡–‡”•ƒ„‘˜‡•‡ƒŽ‡˜‡Ž

ȋ‡–‡ƒͳͻͻ͹Ǣƒ…ƒͳͻͻͻǢ‡…‡”•‡–ƒŽǤʹͲͲͳǢ†‡™‡–ƒŽǤʹͲͲͷǢ ʹͲͲ͸Ǣ••‡ˆƒ

‡– ƒŽǤ ʹͲͳͷ„ȌǤ Ї …”‘’ Šƒ• —‡”‘—• „‡‡ˆ‹…‹ƒŽ –”ƒ‹–• –Šƒ– ƒ”‡ Š‹‰ŠŽ› ’”‡ˆ‡””‡† „› „‘–Š

ˆƒ”‡”•ƒ†…‘•—‡”•ǤЇ•‡–”ƒ‹–•‹…Ž—†‡‹Ȍ‹–•ƒ†ƒ’–ƒ„‹Ž‹–›ƒ†–‘އ”ƒ…‡–‘‡š–”‡‡

‡˜‹”‘‡–ƒŽ …‘†‹–‹‘•ǡ ‹‹Ȍ ‘ •–‘”ƒ‰‡ ’‡•– –Šƒ– ƒˆˆ‡…– •‡‡† ƒ† ‰”ƒ‹ “—ƒŽ‹–› ƒˆ–‡”

Šƒ”˜‡•–ǡ‹‹‹Ȍ—–”‹–‹‘—•†‹‡–”‹…Š‹’”‘–‡‹ǡ•Ž‘™”‡Ž‡ƒ•‹‰…ƒ”„‘Š›†”ƒ–‡ǡ‰Ž—–‡ˆ”‡‡ƒ†Š‹‰Š

‹”‘…‘–‡–ƒ†‹˜Ȍ–Ї•–”ƒ™Šƒ•‰‘‘†“—ƒŽ‹–›ǡ—–”‹–‹‘—•ƒ†’ƒŽƒ–ƒ„އˆ‡‡†ˆ‘”Ž‹˜‡•–‘…

•—…Šƒ•‹Ž‹‰…‘™•ƒ†‘š‡ȋ‡–‡ƒͳͻͻ͹Ǣ —ˆƒ‡–ƒŽǤʹͲͲͳǢ‡œ‡‡Ƭ‡”‹Š—ʹͲͲͳǢ

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ƒˆ–ƒ— ‡– ƒŽǤ ʹͲͳͲǢ ••‡ˆƒ ‡– ƒŽǤ ʹͲͳ͵Ǣ ‡”‡‡™ Ƭ ‡Žƒ— ʹͲͳ͵Ǣ ƒ••ƒŠ— Ƭ ‡„‡™

ʹͲͳ͵Ǣƒ›‡ʹͲͳͶȌǤ‡•‹†‡•–‘„‡‹‰”‹…Š‹…ƒ”„‘Š›†”ƒ–‡ǡ–Ї’”‘–‡‹…‘–‡–‘ˆ–‡ˆˆ”ƒ‰‡•

ˆ”‘ͺΨ–‘ͳͷΨȋ‘ƒ˜‡”ƒ‰‡ͳͲȂͳͳΨȌƒ†—•‡†–‘ƒ‡ƒˆŽƒ–ǡ•‘ˆ–ǡ•’‘‰›ǡ–Š‹„”‡ƒ†

…ƒŽŽ‡†‘Injera’ȋƒ„ƒʹͲͳ͹Ǣ‹”Šƒ—‡–ƒŽǤʹͲͳͺȌǤ•ƒ‰Ž—–‡ˆ”‡‡…”‘’ǡ‹–…ƒ„‡…‘•—‡†

„›’‡‘’އ™‹–Љޗ–‡ƒŽŽ‡”‰›ƒ†Š‡Ž’’”‡˜‡–•—ˆˆ‡”‹‰ˆ”‘…‘‡Ž‹ƒ…ƒ†‘–Ї”†‹•‡ƒ•‡•

…ƒ—•‡†„›Ž‘™–‘އ”ƒ…‡–‘‰Ž—–‡ȋ‹”Šƒ—‡–ƒŽǤʹͲͳͺȌǤ—–”‹–‹‘ƒŽŽ›ǡ–Ї…”‘’•—’’‘”–•͸Ͳ Ȃ͹ͷΨ‘ˆ–Ї–Š‹‘’‹ƒ’‘’—Žƒ–‹‘ȋƒ„ƒʹͲͳ͹Ǣ‹”Šƒ—‡–ƒŽǤʹͲͳͺȌǤ

b. Breeding and agronomy

Breeding

‡ˆˆ™ƒ•ƒ‘‰–Ї‘•–‹’‘”–ƒ–…‡”‡ƒŽ…”‘’•‹–Š‹‘’‹ƒ•‹…‡–Ї‡ƒ”Ž‹‡•–ƒ‰”‹…—Ž–—”ƒŽ

•—”˜‡›…‘†—…–‡†‹ͳͻͶͳȋ‘›…‡ͳͻͶ͵ȌǤЇ…”‘’Šƒ•”‡ƒ‹‡†–Ї‘•–‹’‘”–ƒ–…‡”‡ƒŽ

…”‘’ ‹ –Š‹‘’‹ƒ „‡…ƒ—•‡ ‘ˆ –Ї †‹‡–ƒ”› ƒ† ‡…‘‘‹… ˜ƒŽ—‡• ȋƒ„ƒ ʹͲͳ͹Ǣ ‹”Šƒ— ‡– ƒŽǤ ʹͲͳͺȌǤ‡ˆˆ”‡•‡ƒ”…Š™ƒ••–ƒ”–‡†‹–Š‹‘’‹ƒ‹–ЇͳͻͷͲ•™‹–ЖЇ‘„Œ‡…–‹˜‡‘ˆ‹’”‘˜‹‰

–Ї …”‘’ǯ• ’”‘†—…–‹˜‹–› –Š”‘—‰Š •‡Ž‡…–‹‘ ‘ˆ Ž‘…ƒŽ Žƒ†”ƒ…‡•ǡ ƒ† ”‡Ž‡ƒ•‡† ʹͶ ˜ƒ”‹‡–‹‡•

ȋ‡–‡ƒͳͻͻ͵Ǣ••‡ˆƒ‡–ƒŽǤʹͲͳͳȌǤˆ–‡”ƒ››‡ƒ”•‘ˆ†‹•…‘˜‡”›‘ˆŽ‘™‡”‘’‡‹‰ƒ†

’‘ŽŽ‹ƒ–‹‘ Šƒ„‹–•ǡ –‡ˆˆ Š›„”‹†‹œƒ–‹‘ ™ƒ• „‡‰ƒ ‹ ͳͻ͹Ͷ ȋ‡”Ї ͳͻ͹ͷȌ ƒ† ”‡•—Ž–‡† ‹

†‡˜‡Ž‘’‡– ‘ˆ †‹ˆˆ‡”‡– ˜ƒ”‹‡–‹‡• ‘ˆ –Ї …”‘’Ǥ ”‡‡†‹‰ ‘ˆ –‡ˆˆ Šƒ† ƒŽ•‘ ƒ ‘„Œ‡…–‹˜‡ ‘ˆ

„—‹Ž†‹‰ ‘™Ž‡†‰‡ ‘ Š‘™ –‘ …‘–”‘Ž –Ї ƒ‰”‘‘‹… –”ƒ‹–• ‘ˆ –Ї …”‘’ •—…Š ƒ• Ž‘†‰‹‰ǡ

™Š‹…Š…ƒ…ƒ—•‡ƒ›‹‡Ž†Ž‘••—’–‘ʹͷΨƒ†”‡•—Ž–‡†‹–Ї†‡˜‡Ž‘’‡–ƒ†”‡Ž‡ƒ•‡‘ˆ͵ʹ

˜ƒ”‹‡–‹‡• ȋ‡„‡„‡™ ‡– ƒŽǤ ʹͲͳ͵ȌǤ ƒ”‹‡–‹‡• ‘„–ƒ‹‡† ˆ”‘ –Ї „”‡‡†‹‰ ’”‘…‡•• ‹…”‡ƒ•‡

‰‡‡–‹…‰”ƒ‹›‹‡Ž†‘ˆ–Ї…”‘’‘ƒ˜‡”ƒ‰‡‘ˆͲǤͺ؈”‘–Ї‹”ˆ‹”•–”‡Ž‡ƒ•‡‹ͳͻ͹Ͳ—–‹Žͳͻͻͷ ȋ‡„‡„‡™ ‡– ƒŽǤ ʹͲͳ͵ȌǤ —””‡–Ž›ǡ –Ї”‡ ƒ”‡ Ͷʹ –‡ˆˆ ˜ƒ”‹‡–‹‡• ‹ –Š‹‘’‹ƒ ȋ‡„‡„‡™ ‡– ƒŽǤ ʹͲͳ͵Ǣ‹•‰ƒƒʹͲͳͺȌǤƒ–‹‘ƒŽŽ›ǡ‡„”‡œ‡‹–‰”‹…—Ž–—”ƒŽ‡•‡ƒ”…Ї–‡”ȋœȌ—†‡”

–Ї–Š‹‘’‹ƒ•–‹–—–‡‘ˆ‰”‹…—Ž–—”ƒŽ‡•‡ƒ”…ŠȋȌ…‘‘”†‹ƒ–‡•–‡ˆˆ„”‡‡†‹‰™‹–ЖЇ

‘„Œ‡…–‹˜‡‘ˆ–‡ˆˆƒ‰”‘‘‹…’‡”ˆ‘”ƒ…‡‡Šƒ…‡‡–ǡƒ††‡˜‡Ž‘’Š‹‰Š›‹‡Ž†‹‰ƒ†•–”‡••

–‘އ”ƒ–˜ƒ”‹‡–‹‡•Ǥ

‡‡”ƒŽŽ›ǡ–ЇЋ•–‘”›‘ˆ–‡ˆˆ„”‡‡†‹‰‹–Š‹‘’‹ƒŠƒ•ˆ‘ŽŽ‘™‡†ˆ‹˜‡’Šƒ•‡•ƒ•†‡•…”‹„‡†‹

••‡ˆƒ ‡– ƒŽǤ ȋʹͲͳͳȌǤ Ї ˆ‹”•– ’Šƒ•‡ ȋͳͻͷ͸Ȃͳͻ͹ͶȌ ™ƒ• …Šƒ”ƒ…–‡”‹œ‡† „› ƒ ‡’Šƒ•‹• ‘

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‰‡”’Žƒ• ‡Šƒ…‡‡– ȋ…‘ŽŽ‡…–‹‘Ȁƒ…“—‹•‹–‹‘ǡ …Šƒ”ƒ…–‡”‹œƒ–‹‘ ƒ† ‡˜ƒŽ—ƒ–‹‘ǡ

•›•–‡ƒ–‹…• ƒ† …‘•‡”˜ƒ–‹‘Ȍǡ ‰‡‡–‹… ‹’”‘˜‡‡– ”‡Ž›‹‰ ‡–‹”‡Ž› —’‘ ƒ•• ƒ†Ȁ‘”

’—”‡ǦŽ‹‡•‡Ž‡…–‹‘†‹”‡…–Ž›ˆ”‘–Ї‡š‹•–‹‰‰‡”’Žƒ•ƒ†‹‹–‹ƒ–‹‘‘ˆ‹†—…‡†—–ƒ–‹‘

–‡…А‹“—‡•Ǥ ‡…‘† ’Šƒ•‡ ȋͳͻ͹ͷȂͳͻͻͷȌ ƒ”‡† „› –Ї ‹…‘”’‘”ƒ–‹‘ ‘ˆ ‹–”ƒǦ•’‡…‹ˆ‹…

Š›„”‹†‹œƒ–‹‘‹–‘–ЇƒŽ”‡ƒ†›’”‡Ǧ‡š‹•–‹‰„”‡‡†‹‰‡–Š‘†•ˆ‘ŽŽ‘™‹‰–Ї†‹•…‘˜‡”›‘ˆ–Ї

…Šƒ•‘‰ƒ‘—• ˆŽ‘”ƒŽ ‘’‡‹‰ „‡Šƒ˜‹‘” ‘ˆ –‡ˆˆ ˆŽ‘™‡”• ȋˆ”‘ ƒ„‘—– ͸ǣͶͷȂ͹ǣ͵Ͳ Ȍ ƒ†

–Ї”‡„›–Їƒ”–‹ˆ‹…‹ƒŽ…”‘••‹‰–‡…А‹“—‡ƒ•†‡•…”‹„‡†„›‡”Їȋͳͻ͹ͷȌǤŠ‹”†’Šƒ•‡ǡȋͳͻͻͷȂ ͳͻͻͺȌ ˆ‡ƒ–—”‹‰ ‹‹–‹ƒ–‹‘ ‘ˆ ‘އ…—Žƒ” ƒ’’”‘ƒ…Ї• ‹…Ž—†‹‰ †‡˜‡Ž‘’‡– ‘ˆ ‘އ…—Žƒ”

ƒ”‡”• ƒ† ‰‡‡–‹… Ž‹ƒ‰‡ ƒ’•ǡ ƒ† ƒƒŽ›•‡• ‘ˆ ‘އ…—Žƒ” ‰‡‡–‹… †‹˜‡”•‹–›Ǥ ‘—”–Š

’Šƒ•‡ ȋͳͻͻͺȂʹͲͲ͵Ȍ ƒ”‡† „› ˆ—”–Ї” ‹…‘”’‘”ƒ–‹‘ ‘ˆ ‹ ˜‹–”‘ …—Ž–—”‡ –‡…А‹“—‡• ƒ†

‹–‡”Ǧ•’‡…‹ˆ‹… Š›„”‹†‹œƒ–‹‘ ȋ‡ˆ‡”ƒ ‡– ƒŽǤ ʹͲͲ͵Ȍ ƒŽ‘‰ ™‹–Š ”‡Ǧƒ’’”ƒ‹•ƒŽ ‘ˆ ‹†—…‡†

—–ƒ‰‡‡•‹• ’ƒ”–‹…—Žƒ”Ž› ˆ‘” Ž‘†‰‹‰ ƒ† އƒˆ ”—•– †‹•‡ƒ•‡ ”‡•‹•–ƒ…‡Ǥ ‹ˆ–Š ’Šƒ•‡ ȋˆ”‘

ʹͲͲ͵ –‹ŽŽ ʹͲͲ͸Ȍ ˆ‡ƒ–—”‹‰ ‹–”‘†—…–‹‘ ‘ˆ ’ƒ”–‹…‹’ƒ–‘”› „”‡‡†‹‰ ƒ’’”‘ƒ…Ї• ‹ –Ї ’”‡Ǧ

‡š‹•–‹‰‘˜‡”ƒŽŽ–‡ˆˆ‰‡‡–‹…‹’”‘˜‡‡–˜‡–—”‡•ȋ‡Žƒ›‡–ƒŽǤʹͲͲ͸Ǣ‡Žƒ›‡–ƒŽǤʹͲͲͺȌƒ†

…‘–‹—‡†‡š–‡•‹˜‡‘އ…—Žƒ”ƒ†‰‡‘‹…”‡•‡ƒ”…Šƒ’’”‘ƒ…Ї•–‹ŽŽ’”‡•‡–Ǥ

Š‘—‰Š•—……‡••ˆ—Žǡ–‡ˆˆ„”‡‡†‹‰‡•’‡…‹ƒŽŽ›–Š”‘—‰ŠŠ›„”‹†‹œƒ–‹‘ˆƒ…‡•…ŠƒŽŽ‡‰‡•‘•–Ž›

ˆ”‘–Їƒ–—”ƒŽ„‡Šƒ˜‹‘—”‘ˆ–Ї…”‘’ȋ„‹‘Ž‘‰‹…ƒŽ’”‘„އ•ȌǤƒŽŽ•‡‡†•‹œ‡ǡŽ‘†‰‹‰ǡ•ƒŽŽ

ˆŽ‘™‡”•–Šƒ–‘’‡‘Ž›ƒ–͸ǣͶͷȂ͹ǣ͵Ͳƒ‹–Ї‘”‹‰ȋ‡”Їͳͻ͹ͷȌǡ†‹ˆˆ‹…—Ž–›‘ˆ…”‘••‹‰ǡ

•Šƒ––‡”‹‰ǡ ’”‡ ƒ† ’‘•–ǦŠƒ”˜‡•– ›‹‡Ž† Ž‘••‡•ǡ Ž‘™ ˆ‡”–‹Ž‹œƒ–‹‘ ƒˆ–‡” …”‘••‹‰ ƒ† Ž‹‹–‡†

ˆ‘…—•ƒ”‡ƒ‘‰–Ї…ŠƒŽŽ‡‰‡•‘ˆ–‡ˆˆ„”‡‡†‹‰‹–Š‹‘’‹ƒȋ‡–‡ƒͳͻͻ͵Ǣ‡–‡ƒͳͻͻ͹Ǣ

‡„‡„‡™‡–ƒŽǤʹͲͳ͵Ǣ‡„‡„‡™‡–ƒŽǤʹͲͳ͹Ǣ‹•‰ƒƒʹͲͳͺȌǤ

Agronomy

Cropping season – from the farmer’s perspective

‡ˆˆ‹•…—Ž–‹˜ƒ–‡†‹–™‘•‡ƒ•‘•‡•’‡…‹ƒŽŽ›™Š‡–Ї˜‘Ž—‡ǡ…‘˜‡”ƒ‰‡ƒ††‹•–”‹„—–‹‘‘ˆ

”ƒ‹ˆƒŽŽ‹•‡˜‡ƒ†—‹ˆ‘”ǤЇˆ‹”•–•‡ƒ•‘ˆ‘”–‡ˆˆ’”‘†—…–‹‘‹•ˆ”‘‡†‘ˆ’”‹Ž–‘ƒ›

ˆ‘”•‘™‹‰ƒ†‡†‘ˆ‘˜‡„‡”–‘‡ƒ”Ž›‡…‡„‡”ˆ‘”Šƒ”˜‡•–‹‰ǤŠ‹••‡ƒ•‘‹•Ž‹‹–‡†

–‘ƒ”‡ƒ•™‹–Š™‹†‡”†‹•–”‹„—–‹‘‘ˆ˜‡”–‹…•‘‹Ž•ƒ†‡ƒ”Ž›‘•‡–ƒ†Š‹‰Šƒ‘—–‘ˆ•‡ƒ•‘ƒŽ

”ƒ‹ˆƒŽŽǤЇ•‡…‘†ƒ†–Ї‘•–…‘‘‹•–Ї•‡ƒ•‘™‹–Їƒ”Ž›—Ž›–‘‡ƒ”Ž›—‰—•–ˆ‘”

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•‘™‹‰ ƒ† Žƒ–‡ ‘˜‡„‡” –‘ ‹†Ǧ‡…‡„‡” ˆ‘” Šƒ”˜‡•–‹‰Ǥ ‘•– ‘ˆ –Ї ‹’”‘˜‡† –‡ˆˆ

˜ƒ”‹‡–‹‡• ƒ”‡ †‡˜‡Ž‘’‡† „ƒ•‡† ‘ ƒ† ’”‘†—…‡† †—”‹‰ –Ї •‡…‘† •‡ƒ•‘Ǥ Ї –‹‡ ‘ˆ

•‘™‹‰ƒ†Šƒ”˜‡•–‹‰˜ƒ”‹‡•ˆ”‘‘‡ƒ”‡ƒ–‘ƒ‘–Ї”‹–Š‹‘’‹ƒǤ‘™ƒ†ƒ›•ǡ–Їƒ‹

•—‡”–‹‡‹••Š”‹‹‰ˆ”‘ͷ‘–Š•ȋƒ›Ȃ‡’–‡„‡”Ȍ–‘ˆŽ—…–—ƒ–‹‰ʹȂ͵‘–Š•

ȋ—‡Ȃ—‰—•–Ȍȋ ʹͲͲ͸ǢʹͲͳ͹ƒǢʹͲͳ͹„ȌǤ

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